Mountain Lion

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Puma concolor

Status: Threatened

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Classification: Mammal

Description

The mountain lion—also known as the cougar, puma, panther, or catamount—is a large cat species native to the Americas. Mountain lions are large, tan cats. Their bodies are mainly covered in tawny-beige fur, except for the whitish-gray belly and chest. Black markings decorate the tip of the tail, ears, and around the snout. Mountain lions vary hugely in average body size depending on geographic location—their size is smallest closer to the equator and largest closer to the poles. Generally, though, males weigh between 115 and 220 pounds (52 and 100 kilograms) and females weigh between 64 and 141 pounds (29 and 64 kilograms).

The mountain lion’s range spreads all across the Americas, from the Canadian Yukon to The Strait of Magellan, the greatest of range of any living mammal in the Americas. Mountain lions inhabit a wide range of ecosystems, making their home anywhere there is shelter and prey, including mountains, forests, deserts, and wetlands. They are territorial and have naturally low population densities, which means the species requires large swaths of wilderness habitat to thrive.

Mountain lions are stealthy predators, hunting at night and often lying in wait for prey or silently stalking it before pouncing from behind and delivering a lethal bite to the spinal cord. Typically they prey on deer, but also feed on smaller animals, even insects, when necessary. Like all cats, mountain lions are strict carnivores, and they only rarely consume vegetation.

Life History

Mountain lions can breed year-round. Female mountain lions usually give birth every two years. Litters can range in size from one to six cubs. The young may stay with their mother for as long as 26 months, but usually separate after about 15 months. In the wild, a mountain lion can live up to 10 years. In captivity, they can live up to 21 years.

Conservation

The historic range of the mountain lion included almost all of North and South America. The species was so wide-reaching and populous that it had multiple subspecies that varied based on location. Throughout the 1800s and early 1900s, people feared the mountain lion because it posed a risk to their livestock. The species was maliciously hunted and almost eradicated from the eastern United States. Due to conservation efforts, mountain lion populations in the western United States are stable, although far lower than they were historically. While there are still several thousand mountain lions in the wild, their population has significantly decreased from their historical population due to unsustainable hunting, habitat destruction , and conflicts with livestock.

Mountain lions are an "umbrella species" for conservation because their conservation depends on the preservation of large amounts of habitat. A mountain lion usually requires about 13 times as much area as a black bear or 40 times as much area as a bobcat to thrive. By preserving enough wilderness to support a stable mountain lion population, countless other species of plants and animals that share mountain lion habitat benefit.

The eastern cougar, a subspecies of mountain lion, was declared officially extinct by the U.S. Fish & Wildlife Service in 2011, although individuals from western populations have been confirmed to wander as far as the East Coast. Florida panthers , the other U.S. subspecies of mountain lion, are listed as critically endangered on the endangered species list. There are less than 160 Florida panthers left in the wild.

Learn more about our work through our Save LA Cougars campaign.

Unlike other large cats, they cannot roar. Instead they growl, shriek, hiss and purr, similar to house cats. Mountain lions are territorial and solitary. They use pheromones and physical signs (like claw markings or feces) to define their territory.

Animal Diversity Web, University of Michigan Museum of Zoology

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About Mountain Lions

mountain lion travel range

Biology & Behavior

Mountain lions are known by many names, including cougar, puma, catamount, painter, panther, and many more. They are the most wide-ranging cat species in the world and are found as far north as Canada and as far south as Chile.

Solitary cats, mountain lions are highly adaptable to situations and environments, and this adaptability has enabled them to survive across much of their original range in the America's, despite severe habitat loss and active threats.

While their longitudinal range has remained, their latitudinal range has shrunk by more than half. Mountain lions used to be found throughout the United States, but due to bounty hunts in the early 1900s and threats such as persecution, trophy hunting, poaching, retaliation in response to livestock depredation, kitten orphaning, poisoning and habitat loss and fragmentation, mountain lions are now only found in 15 western states, and the genetically isolated Florida panther remains in the East. For more information about mountain lion life history, evolution and historical range, read Cougar: The American Lion below.

Mountain lions are members of the Puma lineage in the family Felidae. They have a long evolutionary history.

mountain lion travel range

FUN FACT: Cheetahs are also a part of the Puma lineage. Mountain lions and cheetahs are not considered “big cats”, however, they are the largest of the “small cats”. Neither mountain lions nor cheetahs can roar, but both species can purr!

Mountain lions are obligate carnivores, meaning they only eat meat. As such, they have large canines and sharp, specialized carnassial teeth (molars) for shearing and tearing flesh. Mountain lions also have retractable claws, which they use to climb trees and capture prey. Their preferred prey are deer and elk and they have co-evolved with deer and elk as their primary prey.

Mountain lions have extraordinary vision. They have round or oval pupils, which allows them to open as wide as possible at night, but close almost completely in bright light, which protects light-sensitive cells.

Although there is little research on mountain lion hearing, it is known that domestic cats have very acute hearing and can detect prey using both sight and sound. They can hear frequencies in the ultrasonic range and are able to move their rounded ears together or independently to isolate sounds. It is also believed that an enlarged auditory bullae (the portion of the skull surrounding the middle ear) may enhance a cat’s sensitivity to certain sounds.

In mountain lions, the sense of smell is not thought to play a large role in acquiring food and is thought to be more important in social behavior, such as mating.

For more details about mountain lion senses, read Cougar: The American Lion – Chapter 4: An Almost Perfect Predator  below.

Mountain lions are solitary cats. They are most active at dusk and dawn, known as crepuscular.

A relentless hunter. The search for prey is driven by the cat’s hunger and, in the case of a female, the need to feed growing kittens. The hungrier the cat, the greater the tendency to roam, with effort focused on areas where prey was previously found.  The mountain lion navigates its home range in a zigzag course, skirting open areas and taking advantage of available cover.  The cat’s keen senses are focused to pick up the slightest movement, odor, or sound.

For more details about mountain lion behavior, read Cougar: The American Lion – Chapter 4: An Almost Perfect Predator  below.

When is enough… enough?

Mountain lions live a short 13 years in the wild — if they make it to old age. Today, few lions live a full natural lifespan.

It’s a difficult life, full of potentially lethal challenges: even when the lion avoids humans.

They are shot for recreation, for sport, and for trophies. They are shot when a rancher’s livestock is lost, and when pets disappear. They are shot when people are afraid.

Individual lions, and individual people, are diminished whenever we kill a lion unnecessarily. We lose not just a member of the species, but mother lions that might train their kittens to avoid people, large and well-established males that have proved themselves able to survive entirely in the wild, and lions that feel, think, experience and behave in the world as individuals.

Each time we kill a lion, we lose a little bit of what distinguishes us as humans: our capacity for compassion, for making rational decisions that benefit the common good, for overcoming the urge to demonstrate power and dominance at the expense of our neighbors and our environment.

If we lose our big cats, we will mourn a species that we barely understood. Only a few of us will have encountered an individual wild lion.

And this will be the greatest loss: That we knew just enough to save them, enough to change our behavior, enough to make a difference, and that we chose not to act.

mountain lion travel range

Mountain lions are often known for their role as a keystone species, but they are less commonly known as an umbrella species.

While a  keystone species  is a species on which other species in an ecosystem largely depend, such that if it were removed the ecosystem would change drastically, an  umbrella species  is a species that have either large habitat needs or other requirements whose conservation results in many other species being conserved at the ecosystem or landscape level.

Mountain lions play a vital role in not only supporting the health and function of ecosystems on the whole, but also by promoting biodiversity.

Visit our story map to learn more by clicking the image below

mountain lion travel range

Lion Age & Gender

Identification guide.

mountain lion travel range

Cougar: The American Lion

"Mountain lions live a short 13 years in the wild -- if they make it to old age. Today, few lions live a full natural lifespan.

It's a difficult life, full of potentially lethal challenges: even when the lion avoids humans.

They are shot for recreation, for sport, and for trophies. They are shot when a rancher's livestock is lost, and when pets disappear. They are shot when people are afraid.

The cougar works a powerful magic on the human imagination. Perhaps it is envy. This majestic feline personifies strength, movement, grace, stealth, independence, and the wilderness spirit. It wanders enormous tracts of American wilderness at will. It is equally at home in forest, desert, jungle, or swamp. An adult cougar can bring down a full-grown mule deer in seconds. It yields to few creatures, save, bears and humans.

The cougar's solitary and stealthy lifestyle feeds its mystery. Unfortunately, mystery breeds fear, myth, and misinformation. Since our European ancestors first landed on American shores 500 years ago, we have waged war on large predators. The grizzly, wolf, jaguar, and cougar are now gone from the majority of their original ranges, and loss of habitat now looms as the greatest threat to the small populations that survive. Only in the last three decades have wildlife biologists begun to chip away at the fable and folklore and reveal the cougar for the remarkable carnivore that it is.

The Mountain Lion Foundation is one organization encouraging a more enlightened view of our American lion. The Mountain Lion Foundation was instrumental in the passage of the California Wildlife Protection Act of 1990, which banned the sport hunting of cougars in California and set aside $30 million a year for the next 30 years for wildlife habitat conservation. Because of their extensive range and position high in the food chain, saving land for cougars also protects land for other wildlife and plants."

Robert Redford

mountain lion travel range

Cougars were roaming the Americas when humans crossed the Bering land bridge from Asia 40,000 years ago. They watched the Spanish conquer the Aztecs and the pilgrims land at Plymouth Rock. The big cats prowled the banks of the Mississippi, Colorado, and Amazon rivers. They crossed the high, windy passes of the Sierra Nevada, the Rocky Mountains, and the Andes. They witnessed the first Mormons arrive in Utah, prospectors invade the California gold fields, and gauchos herd cattle in Argentina’s pampas. From the Canadian Yukon to the Straits of Magellan – over 110 degrees latitude – and from the Atlantic to the Pacific, the cougar once laid claim to the most extensive range of any land mammal in the Western Hemisphere . (1)

That cougars can be found from sea level to 14,765 feet, (2)  and survive in the dense forests of the Pacific Northwest, the desert Southwest, and Florida’s Everglades, is testimony to the big cat’s resilience and adaptability. These qualities were put to the test as never before when European explorers set foot in the Americas in the 14th century. Early colonists viewed the lion as a threat to livestock, as a competitor for the New World’s abundant game, and most importantly, as the personification of the savage and godless wilderness they meant to cleanse and civilize. Their death sentence pronounced, cougars were hunted, trapped, and shot on sight, and their habitat was stripped away as land was cleared to make way for agriculture and new towns. Today, the lion’s known range in North America has been reduced to areas in the 12 western states, the Canadian provinces of British Columbia and Alberta, and a small remnant population in southern Florida. An increasing frequency of sightings suggest that some populations may still survive in parts of the eastern United States and Canada . (3)

The cougar is listed in dictionaries under more names than any other animal in the world

THE CAT OF MANY NAMES Because of their enormous range, cougars are known as “the cat of many names.” Writer Claude T. Barnes listed 18 native South American, and 25 native North American, and 40 English names . (4)

The cougar is listed in dictionaries under more names than any other animal in the world. . (5)  The Guarani Indians of Brazil called them  cuguacuarana,  which French naturalist Georges Buffon corrupted into  cougar . . (6,7)   Puma  comes from Quichua, an Inca language, meaning “a powerful animal.” . (6,8)  To the Cherokee of the southeastern United States the big cat was  Klandagi,  “Lord of the Forest,” while the Chickasaws called him  Koe-Ishto , or  Ko-Icto , the “Cat of God.” . (7)  In Mexico they are  leopardo , to other Spanish-Americans,  el Leon . . (6,7)

In 1500, Amerigo Vespucci was the first white man to sight and record a cougar in the Western Hemisphere. Vespucci, after whom the two American continents are named, was probing the coastline of Nicaragua when he saw what he described to be lions, probably because of their similarity to the more familiar African lion . (8,9)  Two years later, on his fourth voyage to the New World, Christopher Columbus saw “lions” along the beaches of what are now Honduras and Nicaragua . (6,7,8)  The honor of being the first European to sight a “lion” in North America fell to Alvar Nunez Caheza de Vaca, who in 1513, saw one near the Florida Everglades . (8)

mountain lion travel range

FROM WHENCE CAME CATS? The fossil record of felines is as filled with mystery as today’s cats themselves. Paleontologists and biologists have traditionally relied upon fossils and differences in physical structure of modem animals to map the evolution of a particular species. This has proven difficult with cats for two reasons: most ancestral cats occupied tropical forests, where the conditions for the preservation of fossils is poor; and most of the physical characteristics of cats are related to the capture of prey, with the result that all felines are very similar in structure . (11)  As a result, no less than five different hypotheses have been offered to explain the relationships between the various groups and subgroups of extinct and modern cats . (12)

While there are differing interpretations of the evolution of the cat family (Felidae), a few facts are agreed upon. Modern and extinct carnivores have a common ancestor called  Miacids . These primitive, tree-dwelling carnivores lived in the forests of the Northern Hemisphere 39 to 60 million years ago. Then, about 40 million years ago, a burst of evolution and diversification produced the modern families of carnivores. These new carnivores fall into two major groups: a bear-like group  (arctoids),  consisting of modem bears, seals, dogs, raccoons, pandas, badgers, skunks, weasels, and their relatives; and a cat-like group  (aeluroids),  a lineage including the cats, hyenas (yes, that’s right), genets, civets, and mongooses . (12)

The first cat-like carnivores to appear were the saber- tooth cats, about 35 million years ago . (11)  The sabertooths became extinct about 10,000 years ago worldwide, at the end of the last glaciation . (13)  While the sabertooths met their demise, however, the modem cats were evolving and diversifying. Ancestral pumas lived in North America from three to one million years ago, with modern pumas appearing about 100,000 years ago or less . (11)  The American lion had arrived.

THE FAMILY OF CATS Just as there is disagreement about where cats came from, there is debate over how to classify the 37 species of cats that exist today. I discovered no less than six different proposed classification systems for Felidae during the research for this book. The Latin name  Felis  concolor  was first given to the cougar in 1771 by Carolus Linneaus, the father of taxonomy. (It was Linneaus who devised the binomial system for describing and classifying plants and animals.)

mountain lion travel range

A different approach to the evolutionary and taxonomic puzzle of feline classification was taken recently through the application of the new science of molecular evolution. By examining the rate of change of the genes in the DNA molecules of different cat species. biologist Stephen J. O’Brien and his colleagues revealed that the 37 species of modern cats evolved in three distinct lines. The earliest branch occurred 12 million years ago and includes the seven species of small South American cats (ocelot, jaguarundi, and others). The second branching took place 8 to 20 million years ago and included the domestic cat and five close relatives (Pallas’s cat, sand cat, and others). About 4 to 6 million years ago a third branch split and gave rise to the middle-sized and large cats. The most recent split (1.8 to 3.8 million years ago) divided the lynxes and the large cats. This third line gave rise to 24 of the 37 species of living cats, including the cougar, cheetah, and all big cats . (15)

mountain lion travel range

SUBSPECIES AND STATUS When a species is as broadly distributed as the cougar, regional variations in physical appearance occur. For instance, mountain lions from Alberta look somewhat different than the Florida panther, a fact that relates to the different geographic habitats in which the lion lives . (17)  Wildlife taxonomists recognize these regional variations by dividing  Felis   concolor  into some 26 subspecies or geographic races, scattered across North and South America . (18)  This is similar to the different races or breeds of the domestic dog. Edward A. Goldman, coauthor of the classic,  The Puma: Mysterious American  Cat , explains how the subspecies of cougar are classified: “The subspecies or geographic races of the puma, like those of other animals, are based on combinations of characters, including size, color, and details of cranial [skull] and dental structure(6) Twelve subspecies are recognized north of the border between the United States and Mexico . (6,7,18)  (When writing the scientific name of a particular subspecies, such as the cougar found in Colorado, the subspecies name follows the genus and species. Thus, the Colorado cougar becomes  Felis  concolor hippolestes  or  F.   c. hippolestes.)

The existence and status of the various subspecies of cougars in North America is the subject of heated debate among academics and wildlife professionals.

The existence and status of the various subspecies of cougars in North America is the subject of heated debate among academics and wildlife professionals. The two subspecies found in eastern North America, the eastern panther  (Felis concolor couguar)  and the Florida panther (F.  c. coryi),  are classified as endangered and fully protected . (19)  The Yuma puma  (F.  c.  browni),  a subspecies found along the lower Colorado River, is currently a candidate for listing as endangered . (20)  While cougar populations are considered to be healthy in many parts of western North America, populations adjacent to rapidly expanding urban areas are facing critical habitat loss. In southern California for example, mountain lions in the Santa Monica Mountains and Santa Ana Mountains are fast losing ground to rampant residential development.

APPEARANCE AND SIZE The cougar is plain-colored like the African lion, but is of slighter build with a head that is smaller in proportion to its body. Male pumas do not have the distinctive mane and tufted tail of their Old World cousins . (2)  The absence of a mane led to an early myth about mountain lions: Early Dutch traders in New York were puzzled that the lion skins they obtained were those of females Only. They questioned Indian hunters and were assured that such animals existed, but only in the most inaccessible mountainous places, where it would be foolhardy to attempt to hunt them . (1)

mountain lion travel range

In captivity, cougars have lived as long as 21 years . (24)  In the wild, the cats probably live only half as long. Lack of a reliable way to determine a mountain lion’s age makes exact measurements difficult. In the wild, a 10-year-old lion is likely a very old cat. Experts also disagree over which gender lives longer on the average. Some think the added stress of raising kittens guarantees female lions a shorter life. The lifespan of both sexes in hunted populations is probably shorter . (25)  But, even in the absence of hunting, a short life span is to he expected of a predator that faces the frequent hazards of feeding on prey much larger than itself.

Cougar The American Lion Line Illustrations Copyright (1992-2009) by Linnea Fronce

CHAPTER NOTES

  • Seidensticker J.C. 1991a. Pumas. Pages 130-138 in J. Seidensticker and S. Lumpkins, eds.  Great Cats:   Majestic creatures of the wild . Rodale Press. Emmaus, Pennsylvania.
  • Sunquist, F. C. 1991.  The Living Cats . Pages 28-53 in J. Seidensticker and S. Lumpkins, eds.  Great Cats:   Majestic creatures of the wild . Rodale Press. Emmaus, Pennsylvania.
  • Morse, S. C. 1991. Forest Ecologist and Wildlife Habitat Consultant. Morse and Morse Forestry. Jericho, Vermont. (Personal communication)
  • Barnes, C. T. 1960.  The cougar or mountain lion . Ralton Co., Salt Lake City.
  • Lynch, W. 1989.  The elusive cougar .  Canadian Geographic  August/September: 24-31.
  • Young, S. P., and E. A. Goldman. 1946. The Puma: Mysterious American Cat. American Wildlife Institute, Washington D. C.
  • Tinsley, J. B. 1987.  The puma: Legendary lion of the Americas . Texas Western Press, The University of Texas at El Paso.</st1:placetypew:st=”on”>
  • McMullen, J. P. 1984.  Cry of the panther: Quest for a species . Pineapple Press, Englewood Florida.
  • Guggisberg, C. A. W. 1975.  Wild cats of the world . Taplinger Publishing Co., New York.
  • Hornocker, M. G. 1969b.  Stalking the mountain lion – to save him. National   Geographic  November: 638-655.
  • Kitchener, A. 1991.  The natural history of wild cats . Cornell University Press. Ithaca, New York.
  • Neff, N. A. 1991.  The cats and how they came to be.  Pages 14-23 in J. Seidensticker and S. Lumpkins, eds.  Great   Cats: Majestic creatures of the wild . Rodale Press. Emmaus, Pennsylvania.
  • Van Valkenburgh, B. 1991.  Cats in communities: Past and present.  Page 16 in J. Seidensticker and S. Lumpkins, eds.  Great   Cats: Majestic creatures of the wild . Rodale Press. Emmaus, Pennsylvania.
  • Hornocker, M. G., C. Jonkel, and L. D. Mech. 1979.  Family felidae. Mountain lion  ( Felis concolor) .  Wild Animals of North America . National Geographic, Washington, D.C.
  • O’Brien, S. J. 1991.  Molecular evolution of cats . Page 18 in J. Seidensticker and S. Lumpkins, eds.  Great   Cats: Majestic creatures of the wild . Rodale Press. Emmaus, Pennsylvania.
  • Seidensticker, J. C. 1991b. Introduction to “ The Living cats ” by F. C. Sunquist. Page 28 in J. Seidensticker and S. Lumpkins, eds.  Great   Cats: Majestic creatures of the wild . Rodale Press. Emmaus, Pennsylvania.
  • Shaw, H. 1989.  Soul among lions . Johnson Books. Boulder, Colorado.
  • Anderson, A. E. 1983.  A critical review of literature on puma (Felis concolor ). Colorado Division of Wildlife. Special report Number 54.
  • U. S. Fish and Wildlife Service. 1991.  Endangered and threatened wildlife and plants , 50 CFR 17.11 & 17.12 July 15, 1991. U. S. Government Printing Office: 1991-296-520:50024. Washington D. C.
  • Duke, R., R. Klinger, R. Hopkins, and M. Kutilek. 1987  Yuma  puma (Felis concolor browni ). Feasibility Report Population Status Survey. 22 September 1987. Harvey and Stanley Associates, Inc. Alviso, California. Completed for the Bureau of Reclamation.
  • Currier, M. J. P. 1983.  Felis concolor . Mammalian Species No. 200, pp 1-7. American Society of Mammalogists.
  • Lindzey, F. 1987.  Mountain lion . Pages 656-668 in M. Novak, J. A. Baker, M. E. Obbard, and B. Malloch, eds.  Wild furbearing management and conservation in North America . Ministry of Natural resources, Ontario, Canada.
  • Quigley, H. 1990.  The complete cougar. Wildlife Conservation  March/April: 67.
  • Collette, M. 1991. Founder and President, Wildlife Waystation, Angeles National Forest, California. (Personal communication)
  • Hopkins, R. A. 1991. Wildlife Biologist, H. T. Harvey and Associates. Alviso, California. (Personal communication)

mountain lion travel range

Pregnant females do not prepare elaborate dens. It seems matter that it provides a refuge from predators (coyotes, golden eagles, other cougars) and shields the litter from heavy rain and hot sun. Dens rarely contain any bedding for the young, though a mothers soft belly hair was found in one. (2)-  (This also contradicts the popular misconception, perpetuated largely by some nature movies, that cougars always choose caves as dens.) (6)

Newborn mountain lions enter the world as buff brown balls of fur weighing slightly more than a pound. (1)  Biologists call them kittens or cubs either is correct. Their eyes and ear canals are closed, their coats are covered with blackish brown spots, and their tails are dark-ringed. (2)  This color pattern provides excellent protective camouflage.

Kittens begin nursing within minutes after birth and gain weight rapidly, with males tending to outpace females. (1)  Nursing mothers have eight teats but apparently only six produce milk. Kittens start to compete for nipples the first day and generally suckle the same nipple whenever nursing. (4)  At two weeks of age the kittens’ eyes and ears are opened and they are able to walk. Within 10 to 20 days the kittens may weigh over two pounds. They begin to move awkwardly about, exploring the rock overhang, brushy thicket, or pile of boulders that serves as their den. (5)

While suckling her young the mother must occasionally leave the den to hunt. This is the time of her most restricted movement, because she does not want to venture too far from her vulnerable kittens. Still, she must hunt to sustain herself and replenish her milk. While hunting the female cougar remains within a fixed area called a  home range .  Varying in size from 25 to 400 square miles, (7,8)  home ranges are restricted areas of use in which cougars confine their movements while hunting, searching for a mate, or raising young. Biologists refer to the cougars that occupy home ranges as  residents . Possession of a home range is critically important to a female cougar because it increases her litter’s chances for survival by guaranteeing an established hunting area for the mother.

…kittens learn early to move around their range and not imprint upon a single home site.

By the time kittens are weaned at 2 to 3 months, the mother has moved the litter to one or more additional den sites throughout her home range. This provides greater protection for the young and may be one reason she does not construct elaborate dens. In his book  Soul Among Lions , Arizona cougar specialist Harley Shaw explains that there are other advantages to such behavior: “…kittens learn early to move around their range and not imprint upon a single home site. Home is the entire area of use. Within it, lions are free to move, hunt, and rest as their mood and physiology directs. They are not handicapped the human compulsion to return to a single safe base at night. Home is a large tract of land that they undoubtedly come to know as you and I know the floorplan of our house. They learn to be lions in this home area.” (6)

The physical metamorphosis of young, growing cougars is dramatic, especially their teeth and coat. Teeth are critical to a cougar’s survival, so the teeth in young cougars develop quickly. Their large canines (or fangs) allow them to capture and kill prey, while their specially adapted molars (called carnassials) are used to cut through tissue while feeding. Canines first appear at age 20 to 30 days, followed by the molars at 30 to 50 days. Permanent teeth start replacing primary (baby) teeth at about 5 1/2 months. The permanent canines first appear at month eight, and for a short time both permanent and primary canines are present. (3)

mountain lion travel range

As an adult cougar’s tawny coat provides camouflage while stalking prey, a kitten’s spots provide camouflage from predators. Kittens begin to lose these spots at 12 to 14 weeks, they fade rapidly but are still obvious at 8 months, less so at one year. By 15 months the markings are visible only on the hindquarters and only under certain light conditions. In some cougars, the stripes on the upper foreleg are still visible at 3 years of age. (3,9)

The mountain lion’s coat is not the only feature that changes color with age. Their eyes, light blue at birth, begin to change at four months and are the golden brown of adults by 16 to 17 months. (3,9)

GROWING UP AND LEAVING HOME Female cougars probably begin leading their kittens to kills as early as 7 to 8 weeks. The mother also carries meat to her young from kills until weaning age (2 to 3 months), at which point the cubs weigh in at between 7 and 9 pounds.  As the kittens grow older, the mother will leave them at kills, frequently for days at a time, while she goes in search of the next prey. (6)  As the kittens grow and become stronger, the mother will range farther in search of prey.

Biologists have frequently noted how intensely a female with kittens uses her home range. This is most concentrated subsequent to birth, then expands as the kittens are able to accompany her to kills. It’s easy to imagine an insistent mother as she drags, pushes, and urges her kittens along over the many miles between kills. She expends an enormous amount of energy feeding her growing litter. As a result, the density of prey in the mother’s home range affects how well she can provide for her young, which in turn influences their likelihood of survival.

Arrival at a kill is a time of both feeding and play for kittens. Vegetation is frequently disturbed for 50 feet surrounding the carcass. Grass is flattened, limbs are broken off trees and trunks are covered with the kittens’ claw marks. The carcass is more fully consumed than it would be by an adult lion alone, and pieces of hair and bone are scattered about. This rambunctious play by the young at a kill is another part of their training as predators. They will stalk, attack, and wrestle with their siblings or mother, as if they were the next meal rather than their own flesh and blood. Ultimately, though, play gives way to the real thing.

As they grow stronger and more skilled at stalking, kittens will separate from their mother for days at a time and hunt on their own. This growing independence is a precursor to young lions leaving their mother and going in search their own home range. Biologists are not certain whether a mother and her young gradually grow apart, with the kittens gradually leaving of their own accord, or whether she abandons them as do female black bears with their young. Sonny Bass has found the latter to the case in Florida. “My experience with Florida panthers in the Everglades based on daily tracking) indicates that the mother leaves the young.” (10) Seidensticker tells of one Idaho cougar that abandoned her kittens at a kill. (11)  Paul Beier, who studied mountain lions in southern California, believes the mother discourages her kittens from remaining with her. “Some sort of agonistic behavior on the part of the mother is necessary to discourage the young from staying. Simply abandoning the young is not possible because they know where to find her.” (12)  The presence of mature resident males attracted to the female, who by now is in heat, may also discourage the young from remaining. However they separate, the kittens are finally on their own and the mother will come into heat and breed again. (6)

Kittens can survive on their own as early as 6 months, such as when the mother is killed or dies of natural causes, but this appears to be rare. Typically, the young cougars will remain with their mother for 12 to 18 months. This allows them to hone their hunting skills and gives them time to develop their killing bite. (14)  This bite is usually delivered to the back of the neck of large prey, severing the spinal cord and causing almost immediate death. To be executed efficiently, the bite requires practice and development of the cougar’s powerful jaw muscles. Evidence seems to indicate that the behavioral patterns of killing prey may be innate, but that selection of appropriate prey and stalking may require practice to acquire the necessary skill. (1,2,6)  This may explain why young cougars are sometimes found with a face full of porcupine quills, or are the culprits in attacks on domestic livestock.

The departure of young cats from their mother’s home range is called  dispersal , and it is a time when the young cougars are especially vulnerable; they expose themselves to the dangers of taking prey without the alternative of food provided by their mother. These young cats called  transients,  wander far from the familiar home range of their mother and their hunting skill are not as efficient as those of older resident cats. The dispersal ot young transient cougars out of their birth areas is crucial, however, as it reduces inbreeding and provides new blood to outlying populations. (9)

MATING Both male and female cougars are sexually mature at 24 months, but females have been known to breed as early as 20 months; (9) a Florida panther was recently reported as having given birth before she was 2 years old. (15)  The age of the first breeding may be delayed until the female has established a home range. (16)

When it comes time to mate, the first challenge facing a male and female cougar is finding each other. Solitary and territorial by nature, cougars are frequently scattered over hundreds of miles of rugged terrain. It further complicates the matter that females are receptive to males for only a few days out of each month; (17)  however, it appears to be the lions’ territorial habits and keen senses that ultimately allow them to come together.

Polygamy seems to be the rule for both male and female mountain lions. Males occupy larger home ranges than females, and a resident male with a large home range typically overlaps or encompasses the home ranges of several resident females. Nevertheless, in stable cougar populations with established home ranges, females rarely mate with more than one resident male during a breeding cycle. (9)

mountain lion travel range

Resident male cougars use  scrapes  as visual and olfactory signals to other cougars and to mark their home range area. A scrape (or scratch) is a collection of pine needles. leaves, or dirt scraped into a pile with either the forepaw or hindpaws. Occasionally they urinate or defecate on the pile. Scrapes are made throughout the home range and are frequently located along travelways under a tree (18,19)  or along ridges. Females rarely scrape, more commonly burying their feces under mounds of dirt and debris; these mounds are usually found near large kills. (19)

Mountain lion authority Fred Lindzey believes scrapes help mountain lions both avoid and locate each other. “Scrapes are definitely a means of communication. They broadcast the resident male’s presence to other males (residents and transients) and to females. Females may use scrapes made by the resident male to both avoid him when she has dependent kittens and to find him when she is in estrus.” (20)

Adult males probably spend most of their time searching for receptive females. (21)  When mating does occur, it usually takes place in the female’s home range, with the male seeking out the female. (6)  The female’s estrous cycle lasts approximately 23 days and she is usually in heat for about 8 days. The pair may stay together for up to 3 days, sometimes even sharing a kill. (19)

Cougars compensate for long periods of solitude with some of the most vigorous breeding behavior known to exist among mammals. Copulation can occur at a rate of 50 to 70 times in 24 hours for a 7- to 8-day period. (22)  Each copulation lasts less than a minute. (2)  Such enthusiastic copulation in is thought to stimulate ovulation, (the release of eggs from the ovaries to make them available for fertilization). In his book  The Natural History of Wild Cats , Andrew Kitchener explains the advantage of such behavior: “Most cats are thought to be induced ovulators, so that even though the female may come into estrus, no ovulation occurs unless the vagina and cervix of the female are stimulated repeatedly during mating. As a consequence of estrus lasting several days and ovulation being induced, the chance of a successful fertilization can he maximized.” (23)  Some biologists speculate that high copulation rates also evolved as a way for females to evaluate male vigor (1)  and to ensure that their offspring receive the best genetic endowment. (24)

Cougars appear to be as vocal as they are enthusiastic during mating. The “caterwaul,” characteristic in domestic cats, seems to be even louder in mating cougars. Such behavior has been documented both in captive and wild cougars. (9)  Paul Bier has heard these sounds coming from mating cougars in his California study area; (12)  biologist Susan de Treville, who studied mountain lions in California, was camping on the Malaspina Peninsula in British Columbia when the heard two cougars mating nearby. “Both were screaming loudly. They got to within a foot of my tent, then they gradually moved off. In the morning I found the ground torn up and all the grass flattened.” (25)

After 88 to 96 days, the mother retires to the seclusion of the den and gives birth to a litter of 1 to 6 kittens (or cubs). The average litter size is 2 to 3 kittens, but a young female may produce only 1 kitten in her first litter. This seems to reduce the stress on first-time mothers, allowing them to develop their skills in rearing young. Since cougars tend to bear young every other year, a female that lives for 8 to 10 years has the potential to produce 5 litters. One captive cougar produced 7 litters in 16 years. (27) How many of the kittens survive to adulthood is still a mystery. It is also unknown if the number of offspring produced by a female cougar fluctuates in relation to the abundance of prey, as in other predators such as coyotes and barn owls. Few newly born litters have been studied closely in the wild and definitive information is lacking; however, current research underway in Yellowstone National Park and in the San Andres Mountains of New Mexico may provide some answers about the early lives of pumas.

If a female loses her kittens to predators or other circumstances, she may begin her estrous cycle and breed again soon after the loss. (28)  Sometimes, predators include male cougars; studies in Idaho, Utah, and California have documented that males do indeed kill and even eat kittens on occasion. Whether this is an evolved behavior similar to African lions is unknown, but it may partly explain why females with kittens are unreceptive to males and intolerant of their presence until the young are independent and can hunt for themselves. Females also seem to possess the ability to suppress their estrous cycle during the period they are raising young. Some experts speculate that this ability is hormonal in nature and is possibly related to lactation; others suggest that estrous cycles continue normally and the female simply works harder at avoiding males by being careful where she urinates and by burying her feces. Whether this behavior is hormonal, behavioral, or both is unknown.

Unlike most wild animals, cougars can and do give birth throughout the year, although peaks have been documented in different parts of their range. One population in Idaho peaked in the spring, (16) while cougars in parts of Utah and Wyoming (29) had fall birth peaks. Nevada biologists documented birth peaks during June and July and noted 70 percent of all births occurred between April and September. (9) Mountain lions in and around Yellowstone National Park give birth primarily in midsummer. (30)  Researcher Allen Anderson looked at the birth dates of 6 wild and 35 captive cougars and discovered that over half (55 percent) of the births occurred during April, June, July, and August. (1)

Biologists long speculated that in temperate climates, births occurring during the warmer months placed less stress on both the mother and kittens; however, as Harley Shaw points out, “Birth in warm months forces the mother to be feeding  large  young during mid to late winter. This does not reduce stress on her over the long haul.” (31)  It has also been suggested that in the warmer climates of Arizona, Florida, and California, births may be more evenly distributed throughout the year. Existing information from these states is inconclusive. Two more aspects of the American lion that have left experts scratching their heads.

DEATH While all cougars enter the world in the same fashion, they leave it in a variety of ways. Existing information indicates that the three primary causes of cougar deaths are humans, natural causes, and accidents.

mountain lion travel range

More mountain lions die at the hands of humans than any other known cause of death. This is as true today as it was in the past. A minimum of 65,665 cougars were shot, poisoned, trapped, and snared by bounty hunters, federal hunters, and sport hunters from 1907 to 1978 in the 12 western states, British Columbia, and Alberta. (32)  This carnage seemed to peak between 1930 and 1955, with the highest numbers of pumas killed in California, British Columbia, and Arizona. (1)  This sobering tally does not include the thousands of cougars slaughtered prior to the 1900s nor the untold numbers that have gone unreported since.

Today, cougar hunting is legal in Arizona, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, Texas, Utah, Washington, Wyoming, and the Canadian provinces at British Columbia and Alberta. During the 1989-1990 sport harvest season more than 2,176 cats were killed. (33)  Most of these states allow hunters to kill only one lion per season with the notable exception of Texas, which has the most liberal hunting regulations and places no limits on the number of cats a hunter can take. The cougar enjoys full protection in 24 states and provinces, but has no legal classification and no protection, except in agreement with the Federal Government, in 22 other states and provinces.  (3,32)

Predator control programs present yet another obstacle to the cougar’s survival. The U. S. Department of Agriculture’s Animal Damage Control (ADC) program was responsible for killing 207 cougars in 11 western states during the 1988 fiscal year because of attacks on domestic livestock. (34)

In addition to ADC’s efforts, many states carry on their own predator control programs. For instance, in 1988, ADC killed 38 cougars in California, while the state Department of Fish and Game authorized other hunters to take an additional 28 cougars on depredation permits, for a total of 64 cats. This situation is further complicated by the fact that cougars are occasionally caught in traps set for other animals, and because there is no easy way to release them many are killed. The cats can sometimes pull themselves free of the traps, often at the cost of severed toes or broken bones. Cats that escape with minor injuries may still be capable of taking large prey and surviving, while those with debilitating injuries likely die of starvation. (9)

Collisions with motor vehicles are the primary cause of death in Florida panthers.

Collisions with motor vehicles are the primary cause of death in Florida panthers. From 1979 to 1991, almost 50 percent of documented mortality of the Florida cats was due to collisions with autos. (35)  In California, 22 mountain lions fell victim to collisions between 1971 and 1976, (7)  while researcher Paul Beier lost five lions he was studying to cars. (12)  Three young cougars were even killed by a train, all in the same incident, in Colorado. (1)

A number of the cats have drowned in irrigation canals, (36)  or by falling into wells. (37)  Cougars are capable swimmers, but the smooth concrete banks make escape difficult and the exhausted cats will eventually drown. Unfortunately, such incidents will increase as more cougar habit is encroached upon by humans.

Other types of accidents include falls from cliffs, being struck by lightning, being hit by rock slides, being poisoned by venomous snakes, and choking. (3)  Susan de Treville tells of a mountain lion that died from a violent encounter with a manzanita bush. “We were monitoring an old lion (9-10 years) named Snaggletooth, because he had a broken upper canine. One day we found him lying in an open field-dead. We had no idea what killed him. Later an examination revealed a 5-inch piece of manzanita in the cat’s throat. Apparently, during the final rush at what we think was a deer, the cat ran into a manzanita bush at high speed driving a stab down its throat and severing the carotid artery. Failing eyesight may have been part of the reason Snaggletooth bled to death internally.” (25)

There are three times during their lives when cougars are most at risk: immediately after birth, immediately after becoming independent transients, and during old age. (3) Kittens left alone at a den or kill are vulnerable to other predators, including, as has been noted, adult male cougars; it is unknown how many kittens survive to maturity, but experts suspect that kitten deaths could equal or exceed the number of cougars killed by sport hunting. Transient cougars spend most of their time in unfamiliar territory and have not honed their hunting skills, so do not hunt as efficiently as resident cougars. Old cougars experience extreme tooth wear and loss in weight, making them less efficient hunters, resulting in starvation. Old age is probably the most significant cause of death in unhunted mountain lion populations; a recent study in southern Utah showed that the annual mortality rate in an unhunted cougar population was a fairly high 26 percent.  (26)  In Montana’s hunted cougar populations, over 50 percent of the resident adults in one area were killed, according to research conducted there. (40)

Adult cougars do kill and even eat one another on occasion. (1)  Fighting has been documented in Arizona, California, Nevada, Texas, Wyoming, and Utah. In one study in the San Andres Mountains of southern New Mexico, fighting was found to the primary cause of death. (41)  While in Florida, fighting has led to the death of six endangered panthers over the past 11 years. Of these, two were transient males dispersing from their mother’s home range through home ranges of resident males courting females in heat; two were adult females killed by a young adult male; and the last two were the result of fighting between adult males. (35)  Experts speculate that most conflicts are over females and home ranges, but it is still unknown precisely how much fighting contributes to overall mortality in a cougar population.

Cougars appear to suffer from relatively few internal and external parasites. Those they do contend with include an assortment of fleas, ticks, mites, and tapeworms. The puma’s solitary lifestyle and its habit of spending little time in dens probably minimizes infestation. (2,3)

Deaths attributable to more serious diseases appear to be uncommon. Only two cases of rabies have been documented in wild mountain lions, one in California in 1909, (42)  and a more recent case in Florida. (35)  Naturally occurring antibodies to feline distemper were found in 85 percent of the Florida panthers tested. (43) Another mountain lion in California was recently diagnosed with feline leukemia and was killed. California Department of Fish and Game veterinarian Thierry Work thinks the cat may have been infected by eating domestic cats. The feline leukemia virus is frequently fatal and no vaccine for wild cougars exists; this disease especially threatens small, isolated populations of cougars that front on urban areas, such as in southern Florida and southern California. Allen Anderson cautions that the widely held opinion that wild pumas are largely free of parasites and diseases may be due to the lack of specific research rather than reality. (1)  Cougar diseases are just one of many aspects of the cat that need further study.

  • Anderson, A.E. 1983.  A critical review of literature on puma (Felis concolor) . Colorado Division of Wildlife. Special Report Number 54.
  • Dixon, K.R. 1982. Mountain lion. Pages 711-727 in J.A. Chapman and G.A. Feldhamer, eds.  Wild mammals of North America . John Hopkins University Press. Baltimore.
  • Currier, M.J.P. 1983.  Felis concolor . Mammalian Species No. 200, pp. 1-7. American Society of Mammalogists.
  • Eaton, R.L. and K.A. Velander. 1977. Reproduction in the puma: Biology, behavior and ontogeny. Pages 45-70 in R.L. Eaton, ed.  The world’s cats , Vol. 3(3):  Contributions to breeding biology, behavior and husbandry . Carnivore Research Institute, University of Washington, Seattle.
  • Young, S.P., and E.A. Goldman. 1946.  The puma: Mysterious American cat . American Wildlife Institute, Washington, D.C.
  • Shaw, H. 1989.  Soul among lions . Johnson Books. Boulder, Colorado.
  • Sitton, L.W. and S. Wallen. 1976.  California   mountain lion study . California Department of Fish and Game. Sacramento.
  • Hemker, T.P., F.G. Lindzey, and B.B. Ackerman. 1984. Population characteristics and movement patterns of cougars in southern Utah.  Journal of Wildlife Management , 48(4):1275-1284.
  • Lindzey, F. 1987. Mountain lion. Pages 656-668 in M. Novak, J.A. Baker, M.E. Obbard, and B. Malloch, eds.  Wild furbearer management and conservation in North America . Ministry of Natural Resources, Ontario, Canada.
  • Bass, O.L. 1991. Wildlife Biologist, Everglades National Park Research Center, Homestead, Florida. (Personal communication)
  • Turbak, G. and A. Carey. 1986.  America’s great cats . Northland Publishing, Flagstaff, Arizona.
  • Beier, P. 1992b. Project Leader, Orange County Cooperative Mountain Lion Study, Department of Forestry and Resource Management, University of California, Berkeley. (Personal communication)
  • Hornocker, M.G. and G.M. Koehler. 1985. Reintroducing orphaned mountain lion kittens into the wild. Pages 167-169 in J. Roberson and F. Lindzey, eds.  Proceedings of the second mountain lion workshop , Salt Lake City.
  • Bogue, G. and M. Ferrari. 1974. The predatory “training” of captive reared pumas. Pages 36-45 in R.L. Eaton, ed.  The world’s cats, Vol. 3( 1): Contributions to status, management and conservation . Carnivore Research Institute, University of Washington, Seattle. (Cited from Dixon 1982.)
  • Maehr, D.S., J.C. Roof, E.D. Land, and J.W. McCown. 1989. First reproduction of a panther ( Felis concolor coryi ) in southwestern Florida.  Mammalia , 53:129-131.
  • Seidensticker, J.C., IV, M.G. Hornocker, W.V. Wiles, and J.P. Messick. 1973. Mountain lion social organization in the Idaho Primitive Area.  Wildlife Monographs , 35.
  • Rabb, G.G. 1959. Reproductive and vocal behavior in captive pumas.  Journal of Mammalogy . 49:616-617. (Cited from Dixon1982.)
  • Shaw, H. 1987.  Mountain lion field guide . 3rd Edition. Special Report Number 9. Arizona Game and Fish Department.
  • Hemker, T.P. 1982. Population characteristics and movement patterns of cougars in southern Utah. M.S. thesis, Utah State University, Logan.
  • Lindzey, F.G. 1991. Wildlife Biologist, U.S. Fish and Wildlife Service, Wyoming Cooperative Fish and Wildlife Research Unit, University of Wyoming, Laramie, Wyoming. (Personal communication)
  • Hopkins, R.A. 1991. Wildlife Biologist, H.T. Harvey and Associates, Alviso, California. (Personal communication)
  • Eaton, R.L. 1976. Why some felids copulate so much.  World’s cats . 3:73-94. (Cited from Anderson1983.)
  • Kitchener, A. 1991.  The natural history of the wild cats . Cornell University Press. Ithaca, New York.
  • Lynch, W. 1989. The elusive cougar.  Canadian Geographic  August/September: 24-31.
  • de Treville, S. 1991. Wildlife Biologist. de Treville Environmental Engineering. San Diego, California. (Personal communication)
  • Lindzey, F.G., B.B. Ackerman, D. Barnhurst, T. Becker, T.P. Hemker, S.P. Laing, C. Mecham, and W.D. Van Sickle. 1989.  Boulder   Escalante cougar project final report . UtahDivision of Wildlife Resources, Salt Lake City, Utah.
  • Conklin, W.A. 1884.  The mammals of the Adirondack Region, northeastern New York . L.S. Foster Press, New York. (Cited from Dixon 1982.)
  • Hornocker, M.G. 1970. An analysis of mountain lion predation upon mule deer and elk in the Idaho Primitive Area.  Wildlife Monographs . 21:1-39.
  • Logan, K.A. 1983. Mountain lion population and habitat characteristics in the Big Horn Mountains of Wyoming. M.S. thesis, University of Wyoming, Laramie.
  • Murphy, K. 1991. Wildlife Biologist. Wildlife Research Institute, Inc., Moscow, Idaho. (Personal communication)
  • Shaw, H. 1991. Wildlife Biologist, General Wildlife Services, Chino Valley, Arizona. (Personal communication)
  • Nowak, R.M. 1976.  The cougar in the United States and Canada . New York Zoological Society and U.S. Fish and Wildlife Service Office of Endangered Species, Washington, D.C.
  • Tully, R.J. 1991.  Summary of 1991 questionnaire on mountain lion hunting regulations . Mountain Lion-Human Interaction Symposium and Workshop, April 24-26, Denver. Colorado Division of Wildlife.
  • U.S. Department of Agriculture. 1990. Animal and Plant Health Inspection Service.  Animal damage control program, draft environmental impact statement-1990 .
  • Maehr, D.S., E.D. Land, and M.E. Roelke. 1991b. Mortality patterns of panthers in southwest Florida.  Proceedings of the annual conference of southeast fish and wildlife agencies . 45:In press.
  • Macgregor, W.G. 1976. The status of the puma in California. Pages 28-35 in R.L. Eaton, ed.  The world’s cats, Vol. 3(1): Contributions to status, management and conservation . Carnivore Research Institute, University of Washington, Seattle. (Cited from Dixon 1982.)
  • Sitton, L.W. and R.A. Weaver. 1977.  California   mountain lion investigations with recommendations for management . California Department of Fish and Game, Sacramento.
  • Seidensticker, J.C. 1991a. Pumas. Pages 130-138 in J. Seidensticker and S. Lumpkin, eds.  Great cats: Majestic creatures of the wild . Rodale Press. Emmaus, Pennsylvania.
  • Lopez, B. 1981. The elusive mountain lion.  GEO  June: 98-116.
  • Murphy, K. 1983.  Characteristics of a hunted population of mountain lions in western Montana. (Relationships between a mountain lion population and hunting pressure in western Montana.)  Report to the Montana Department of Fish, Wildlife and Parks.
  • Logan, K.A. 1991. Wildlife Research Institute, Inc., Moscow, Idaho. (Personal communication)
  • Storer, T.I. 1923. Rabies in a mountain lion.  California   Fish and Game . April 9(2):45-48.
  • Roelke, M.E. 1987.  Florida   panther biomedical investigation. Annual performance report . Endangered Species Project E-1-11. Florida Game and Fresh Water Fish Commission. (Cited from Belden 1989.)

mountain lion travel range

WHERE COUGARS LIVE Biologists marvel at the cougar’s remarkable adaptability. The best example of this is the cat’s enormous geographic range. Cougars seem equally at home in Alberta’s alpine forests, Arizona’s Sonoran Desert, or Mexico’s tropical jungles. While the lions don’t seem particular about where they live, studies show that the cats do prefer certain types of terrain and vegetation. Habitat, a space and an environment suited to a particular species, and geographic range, a broader term indicating the map area in which a species occurs, are important concepts in understanding cougar life. (1)  One expects to find cougars only in suitable habitats within a geographic range, and suitable cougar habitat contains two elements: cover and large prey.

Mountain lions are stalking predators that must get close to their prey before ambushing from a short distance. They will take advantage of terrain (steep canyons, rock outcroppings, boulders) or vegetation (dense brush, thickets) to remain hidden while stalking.  Cover  refers to this combination of terrain and vegetation that allows the cat to stay out of sight while hunting and stalking; habitats that have good  stalking cover  attract mountain lions. Cover also helps protect the female cougar’s vulnerable kittens. (In  The Cycle of Life  Chapter I explained how a dense thicket or pile of boulders is used as a den. This is what we mean by  protective cover .)

Even the best stalking cover is of no value, however, if there is nothing to stalk. Deer are the lion’s primary prey-mule deer in western North America and white tailed deer in eastern North America (2)- and deer must be present in sufficient numbers in the lion’s habitat for the cat to survive. (3)  While lions will take a variety of prey, killing one deer is more energy-efficient than killing several rabbits or squirrels, allowing the cat to procure a lot of fresh meat at once. This is particularly important to females with hungry kittens to feed. Since the presence of cougars in an area is largely dependent on the presence of deer, it is important to understand what makes up good deer habitat.

Like cougar habitat, suitable deer habitat must contain a combination of cover and food. But because deer are ungulates (hoofed, plant-eating mammals) they use both resources differently. Where the puma requires stalking cover, deer need  escape   cover,  usually a dense tangle of vegetation into which predators cannot easily follow. (1)  In the case of desert bighorn sheep ( Ovis canadensis),  on which cougars occasionally prey in New Mexico, Nevada and California, escape cover may not be “cover” in the normal sense, but a sheer rock face. The steep incline and exposure discourages a cougar from following and allows the sheep to escape.

Other types of cover are required. Deer need cover for giving birth to fawns and for resting  (protective cover  and  resting cover ). They cope with seasonal temperature extremes by taking refuge in timber stands  (thermal cover ); tall trees with dense foliage reduce heat loss through radiation on cold, clear winter nights, provide shade on hot sunny days, and serve as shields from strong winds for deer and cougars alike. (4)

Being herbivores (plant-eaters), deer will gravitate to habitats that have adequate forage. Mule deer  (Odocoileus hemionus ), the most common species of deer in the western United States and Canada, require a mix of food types (5)  and are known to eat 788 plant species. (6)  In southern Utah, mule deer feed extensively on bitterbrush and Gambel oak, two plants that also provide excellent cover. Not surprisingly, areas dominated by these two plants are also frequented by mountain lions. (7)

Maurice Hornocker (8)  and later John Seidensticker and his coworkers studied cougars in the remote wilderness of the Idaho Primitive Area (now the Frank Church River of No Return Wilderness). There, they found that cougars referred steep, rocks areas covered with dense stands of Douglas fir and ponderosa pine, with sagebrush and grasslands mixed among the bluffs and talus slopes. The big cats avoided crossing large open areas with insufficient cover, preferring to travel around the perimeters.

Researchers Kenny Logan and Larry Irwin (10)  studied cougars in the Bighorn Mountains of northern Wyoming; their work provided the first quantified evaluation of cougar habitat use, and their findings were similar to those of Seidensticker and Hornocker in Idaho. The cats frequented canyonland habitats with steep, rugged slopes (greater than 45 degrees) containing mixed conifer and brushy mountain mahogany cover. Grasslands and sagebrush areas with gentle slopes (less than 20 degrees) were generally avoided.

mountain lion travel range

The late Steven Laing was a member of a team of researchers who studied cougars in the Boulder-Escalante region of south-central Utah. This 10-year study, completed in 1989, is considered to be the most thorough examination of the cats yet completed. Laing, being responsible for examining cougar habitat use, found that cougars selected pinyon-juniper woodlands with lava boulders scattered in the understory cover (ground vegetation). The cats also frequented ponderosa pine/oak brush, mixed aspen/spruce-fir, and spruce-fir habitats. They avoided sagebrush bottomlands, agricultural and pasture lands, slickrock sandstone canyons, and open meadows. These areas were typically at higher elevations with steeper slopes and denser understory cover. This combination of terrain and vegetation seems to enhance the cat’s ability to survey and move through the landscape unseen. (7)

Harley Shaw observed a similar pattern in Arizona. Forested areas such as those found on the Mogollon Rim and Kaihab Plateau have little understory cover and hold relatively, low mountain lion densities, while chaparral and pinyon-juniper vegetations, which have dense understory cover, have higher densities of mountain lions. Shaw suspects understory cover for stalking is the key to habitat suitability. (11)

Cougars cannot successfully take prey if there is too much or too little cover.

Seidensticker and his colleagues (9)  described suitable cougar habitat as a combination of vegetation, topography, prey numbers, and prey vulnerability. Prey vulnerability refers to the ease a prey species may be captured and killed by cougars and depends on the availability of cover (both stalking cover and escape cover) and the behavior of the prey. The relationship between vegetation and prey vulnerability is particularly important: cougars cannot successfully take prey if there is too much or too little cover. The best vegetation for stalking cover is moderately dense-thick enough for the lion to remain hidden, sparse enough for the cat to see its prey. (12)  Thus, the presence of moderately dense stalking cover in a habitat increases the vulnerability of the prey found there. This also explains why dense vegetation is attractive to deer as escape cover-it reduces their vulnerability to cougars.

Writer Barry Lopez ( a MLF Honorary Boardmember ) calls the cougar “a dweller on the edge.” Edges, or  ecotones , are transitional borders between different habitat types-the places where forest meets clearing, where rocky ledge meets bush, and where willow thicket meets streamside banks. (2,13)  Such areas provide good forage and cover for deer, which in turn attracts cougars. Florida panthers make frequent use of ecotones in Everglades National Park; (14)  much of the panthers’ habitat in that part of south Florida is slash pine woodland s with dense understory cover of saw palmetto. This woodland forms the eastern boundary of a flat, grassy wetland (the Everglades) dotted with islands of hardwood trees called hammocks. Panthers do most of their hunting and make most of their kills along the edges of these hammocks and in these woodlands. Laing found a similar pattern in the Boulder-Escalante study: “Riparian zones [streamside habitats] and rock ledges were the two ecotones most associated with highly used areas suggesting habitat selection based on prey densities, cover diversity, and possibly water availability.”  (7)

To a cougar, vegetation shape and density seems to be more important than vegetation type in determining habitat suitability. This partially explains the cat’s ability to occupy such an extensive range and variety of habitats. (12)

So the best deer and cougar habitats appear to be forested areas that contain good deer forage and cover as well as a diversity of terrain and sufficient stalking cover.  Cougars seem consciously to select cover and terrain that allow them find prey, observe it while stalking, and approach close enough to make a kill. (4)  Seidensticker’s study in Idaho (9)  showed that, over a span of years, kill sites were clustered in certain areas, which suggested that these areas offered advantages in taking prey.

HOME RANGE Within mountain lion habitat, adult cougars space themselves out and confine their movement to individual fixed areas known as home ranges. Home range should not be confused with geographic range, which is a broader term indicating the entire map area in which cougars occur. (1)  Cougar home ranges include hunting areas, water sources, resting areas, lookout positions, and denning sites where kittens or cubs can be safely reared. Cougars that occupy home ranges are called  residents , and possession of a home range enhances a resident’ lion’s chances of more consistently finding prey, locating mates, and successfully rearing young. (15)

mountain lion travel range

Possession of a home range is fundamental to the cougar’s survival as a solitary predator. By having a fixed area of land to hunt in, the cougar is better able to consistently locate prey. It roams its home range constantly, learning the terrain, where the best cover is, where the deer most likely can be found. This is why survival for a transient cougar is more precarious than that of a resident cougar.  Transients are constantly moving through unfamiliar territory and have not yet perfected their hunting skills.

Cougars are not territorial in the sense that they defend their home ranges to exclude all other cougars. Rather, the big cats have evolved a  land tenure  system (9)  in which home ranges are maintained by resident lions but not transient lions. Male home ranges are typically larger than female home ranges, usually overlapping or encompassing several of the female ranges, but only occasionally overlapping those of other resident males; however, female home ranges commonly overlap. Exceptions to this pattern do exist. Studies in the Diablo Mountains of California (16)  and the San Andres Mountains of New Mexico (17)  showed overlap between male home ranges, while those of females did not.

In areas where home ranges overlap, cougars seem to avoid each other.

In areas where home ranges overlap, cougars seem to avoid each other. This  mutual   avoidance  is thought to be accomplished primarily through sight and smell. (2, 12)  Smell is employed through the use of scrapes. In the “ Cycle of Life ” chapter it was explained how scrapes function as biological traffic signals within home ranges. By either making a scrape or sniffing the scrape of other individuals, cougars send and receive a variety of messages. Male residents can announce their presence, transient lions or females with dependent kittens can avoid male residents, and females can find males when they are ready to mate. (18)  Two cougars in overlapping home ranges can both use the common area because scrapes allow them to use the area at different times.

After sniffing a scrape impregnated with the urine of another cougar, a lion will display a lip-curling grimace known as a  flehman.  This action is thought to allow them to use a special olfactory organ in the roof of their mouth to evaluate, or get a better “look” at the scent. Biologists speculate that males may use the flehman to determine from a female’s urine whether she is ready to mate. (9,19,20)  Both Seidensticker (9)  and Lindzey (18)  have observed this behavior in captive mountain lions.

mountain lion travel range

Experts speculate that land tenure and mutual avoidance allow cougars too maintain home ranges in a number of beneficial ways. First they seem to reduce conflict. A large, powerful carnivore like the cougar depends exclusively on its good health to capture prey. Frequent fighting could lead to serious injury and starvation. This is not to say fighting between cats never occurs; as has already been pointed out, in some cougar populations, such as in southern New Mexico, fighting is common and even a major source of mortality. (17)  Secondly, maintaining a matrix of adjacent and overlapping home ranges seems to limit cougar population density, (8,21,22)  which in turn increases the cats’ chances of finding prey. Finally, because home ranges are frequently hundreds of square miles in size, it would be impossible for a cougar to actively defend the entire area against all intruders. (15)  As a result, a more flexible system of coexistence has evolved.

Researchers have learned much of what they know about home ranges through radio telemetry. This involves capturing a wild cougar and attaching a collar containing a small radio transmitter. By plotting the cat’s locations on a topographic map, biologists can learn the size of the area a cougar uses throughout the year, what the density of cats in an area is, and the social structure of the population. (23)  What they have learned is that home ranges vary widely in size, depending on local vegetation, prey density, and the time of the year. Male home range size can vary from 25 to 500 square miles, while females usually occupy smaller areas of from 8 to over 400 square miles. Sitton and Wallen documented some of the smallest home ranges in the Big Sur region of coastal California, where the average home ranges were 25 to 35 square miles for males and 18 to 25 square miles for females. (24)  A warmer climate and abundant forage probably make it unnecessary for the deer to migrate between winter and summer range and the herds thus concentrate in specific areas. Consequently, the cougar population concentrates around the denser prey population. The presence of good stalking cover is likely a factor as well. Hemker and his colleagues found some of the largest home ranges during the Boulder-Escalante study in southern Utah, where males occupied areas of up to 513 square miles and females up to 426 square miles. (25)

As might be expected, male and female cougars use their home ranges differently.

As might be expected, male and female cougars use their home ranges differently. Besides a place to hunt, males require in area where they can mate with as many females as possible without interference from surrounding males. That is why male home ranges overlap two, three, or more female home ranges and why male home ranges usually do not overlap. Female home ranges are usually smaller and are used to provide sufficient prey and denning sites for rearing kittens, even in years of low prey density. (15)

Life is tough for a female cougar-much tougher than for a male. Seidensticker explains why: “Adult females are subject to more stress and hazards than are males. The female must hunt and kill large, potentially dangerous prey more frequently than males and must do so at regular, predictable intervals if she is to succeed in rearing her kittens, thus increasing the likelihood of accidental death.” (9)  On occasion, males have even been known to abandon their home range and move to a new area; (26)  older males can also be displaced from their home ranges by prime males. (17)  Females show much more attachment to their home ranges and tend to remain in the same area for their entire lives.

mountain lion travel range

As a result of the functions of land tenure and mutual avoidance, cougars appear to “saturate” an area at a given density. Research indicates that the density of cougars in a particular area is socially regulated through home range size and overlap and does not increase above a level socially tolerable for cougars. In other words, healthy cougar populations appear to be self-regulating. Harley Shaw explains: “In all studies of lions where relatively good documentation of lion numbers has been made, lion densities have peaked and stabilized at points between ten and twenty square miles per adult resident. Evidence indicates that, if left alone, adult resident lions will probably not populate beyond such densities.” (22)

POPULATION DYNAMICS Wildlife biologists have long puzzled over the cougar’s solitary life style and how it benefits the animal as a predator. “I find it curious that an animal starting in a litter becomes antisocial in adulthood,” writes Harley Shaw. “The earliest moments of a lion’s life involve touching its siblings. Its growth involves play, interaction, and cooperation during early efforts in hunting. Yet the animal ultimately comes to avoid other adults except at breeding time.” (22)

The cooperative hunting methods of wolves have been extensively studied and their success is well known, as is their gregarious life style. One cannot help but wonder if in comparison the cougar’s solitary life style, especially hunting, puts the cat at a distinct disadvantage. Seidensticker and his fellow researchers don’t think so: “The mountain lion, too, kills large potentially dangerous prey, but unlike the wolf, a pursuit predator, the lion is a stalking predator whose success depends solely on the element of surprise. In the broken land where lions find sufficient cover to stalk and launch successful attacks, the prey usually are scattered and time-consuming to find. Under such conditions, a solitary social structure is apparently the most effective life style.” (9)

mountain lion travel range

While hunting alone may have its advantages, cougars may not be as solitary as once thought. Rick Hopkins points out that adult males are not solitary by choice and probably spend most of their time searching for receptive females. (27)  Susan de Treville once monitored four radio collared adult resident males to within 50 yards of each other in the Big Sur region of California. (30)  Andrew Kitchener writes: “…far from having a chaotic, random system of home ranges driven by the need for [solitary] cats to avoid each other at all costs, most wild cats maintain a predictable system of land tenure, which promotes social stability and maximizes the reproductive success of both males and females… far from being strangers, neighboring cats probably know each other very well from their own distinctive smells.” (15)

Solitary or not, cougars that inhabit a common geographic area are referred to as a  population , (1)  and within a population not all cougars are created equal. This feline social hierarchy consists of three classes of animals: resident adult males and females, transient males and females, and dependent offspring of resident females. Resident adults maintain established home ranges and do most of the breeding in a population. Transients constantly move through the home ranges of residents in search of a vacant home range of their own; while female transients tend to delay breeding until they find and occupy a home range, this may not be true of males. Dependent offspring include kittens and juveniles that still rely on their mother to hunt for them. (23)

Determining whether a cougar is an adult, transient, or kitten is more complex than it sounds; finding a reliable way to determine age is something which has long eluded biologists. Age is usually established using a combination of tooth wear, body weight, color spotting, and behavior. Kittens are newborn to 16 months old and are still with their mother in her home range; they may still have spots which fade by the third of fourth month. Transients are 17 to 23 months old and have left their mother’s home range but have not yet settled in a home range of their own; spotting may still be present on the insides of the front legs. Resident adults are mature cats, at least 24 months old, and occupy an established home range; spotting is absent or very faint, and females may show evidence of nursing. (26)

mountain lion travel range

Researchers Kenny Logan and Linda Sweanor prefer the terms emigrant and disperser to transient. These refer to lions that have emigrated or dispersed from their birth area but have not set up a home range. This dispersal out of a previously occupied area is called  emigration , while movement into a new area is called  immigration . (1)  Emigrants become immigrants when they enter a new population. (17)  (While it is good science for researchers to refine their techniques and terms, the lack of consistency also makes it difficult for scientists to compare information. It also frustrates writers attempting to explain mountain lions.)

Because no reliable method yet exists for accurately aging cougars, determining how many resident adults, transients, and kittens are present in a cougar population at any given time is equally difficult. This is further complicated by the fact that young transient cats are constantly on the move. (12)  These factors, if not accounted for, can distort population estimates made by wildlife biologists.

Resident females generally outnumber resident males in a population. (25,32,33)  This may be because females require smaller home ranges that frequently overlap those of other female cougars. Females also seem generally more tolerant of adjacent females and, as a result, more females might concentrate in a smaller area. Resident males, on the other hand, require larger home ranges that rarely overlap, and they tend to be intolerant of other males and are more widely scattered than resident females.

mountain lion travel range

Young transient cougars usually leave their mother’s home range (disperse) sometime during their second year. The newly independent cats occasionally linger in the birth area, while others leave immediately. They may wander for more than a before establishing their own home range. (22)  If sufficient space is available, however some females will remain in their birth areas; one young female in Nevada stayed in the mountain range of her birth, bred at 24 months, and established a home range adjacent to her mother’s. Female transients have even been known to take over their mother’s home range after her death. (18)

Males tend to wander farther than females, (26)  although how far either gender travels seems to vary. Transient males in the Diablo Mountains of California traveled up to 35 miles from their areas of birth. (16)  In one Nevada population, males covered an average of 31 miles, while females averaged 18 miles. (26)  One young cougar marked in the Bighorn Mountains of Wyoming appeared in northern Colorado, 300 miles from the original location. (34)

Transient males and females seldom remain in one location longer than six months.

Transient males and females seldom remain in one location longer than six months. How the cats “test” a population for vacant home ranges is poorly understood. It is generally thought that a transient either finds a vacant home range or displaces an older resident. If a vacant home range is found, the young male or female will settle down to the business of establishing its place in the population. If the population density is so high that no vacant home areas exist, the transient moves on.

mountain lion travel range

Kenny Logan believes male transients enter a new population and for a period of time, make kills, avoid other lions, and avoid scrapings. If they think they can establish a home range they begin to scrape. Meanwhile, the resident male, closely monitoring its home range, is busy investigating the newcomer’s scrapes and kills. If the resident and newcomer encounter each other they may fight. As a result, the resident may kill the newcomer, the newcomer may kill the resident, or the newcomer may drive off or displace the resident. (17)

The role of transients in populations is important because they are the primary source of replacements for resident cougars who die as a result of hunting, old age, or accidents. Transients also ensure genetic mixing between populations, (12)  and appear to be a major factor in the recovery of hunted populations. Cougar populations in isolated mountain ranges, such as those common to the Great Basin, are particularly vulnerable because immigration of transients is low. (35)

For example, aggressive predator control efforts wiped out all cougars in and around Yellowstone National Park by the 1920s, and – except for a few transients passing through – no cougars lived there for almost 50 years. But transients seem to be slowly recolonizing the 2.2 million-acre park; research biologist Kerry Murphy directs a field study of cougars in the region and estimates 14 to 17 resident mountain lions are now present. Radio collars have been attached to 9 females and 7 males, and the population produced 21 kittens in 1991. The region has abundant deer and elk, and the cougar population is increasing. (36)

As mentioned earlier, the availability of good hunting sites and prey numbers limit the number of cougars in an area. But birth, death, emigration, and immigration are all factors that greatly influence cougar population maintenance and growth. In absence of hunting by humans, the cougar population will remain relatively constant. (12)  Rick Hopkins found that the unhunted cougar population in the Diablo Mountains of California was relatively stable with a low turnover of residents. (16)

COUGARS ON THE MOVE There is nothing more distinctive about cats than the way they move, and mountain lions are no different. The big cats are the epitome of graceful, lithe motion. Stealthy shadows that do not move so much as flow across the landscape.

Consider the effect of broken terrain on prey, explains Kenny Logan. “Mountain lions are ballerinas at getting across broken terrain-steep slopes, boulders, outcroppings, and undercut ledges.” Mule deer and elk are not as good at negotiating such rugged landscapes. (17)  Here the cougar uses both cover and superior agility to its advantage, and employs yet another weapon in its predatory arsenal-it hunts under the cover of darkness.

Cougars are not strictly nocturnal, as many once thought.

Cougars are not strictly nocturnal, as many once thought. Rather, they tend to be active at the same time as their prey, and deer tend to be active at dawn, dusk, and at night. Animals that are active during the twilight of dawn and dusk are said to be  crepuscular.  The big cat’s excellent night vision makes it well suited for stalking during these low light periods. Florida panthers appear to be more active at sunrise and sunset, (37)  and Paul Beier reports that in southern California mountain lions are active throughout the night. (39)  During the winter in Idaho, radio-collared cougars were found to be more active at night (40 percent of the time they were located) than during daylight hours (14 percent of the time they were located). Daytime activity increased during the summer with an increased abundance of ground squirrels. (9)  Bruce Ackerman studied cougar activity patterns in southern Utah and found the cats most active at sunrise and sunset, but less so during the winter. He also found that single adults were more active than females with young, but that the females became more active as their kittens grew. (3)

Most mule deer herds in mountainous areas of western North America migrate to lower elevations in the winter to avoid heavy snow and then come back to higher elevations in the summer. Predictably, cougars also shift their home range use to follow these seasonal movements. (25)  This elevational movement of cats following deer herds has been observed in Arizona, Idaho, California, and Nevada; in Idaho, cougars’ winter home ranges were smaller than summer home ranges. (9,25)  Biologist Dave Ashman and his coworkers documented a case in Nevada where a male cougar occupied one mountain range in winter, then crossed 10 miles of flat, open desert to take up summer residence in another mountain range. Elsewhere in the state, cougars followed deer herds to lower elevations or traditional wintering grounds. During the cold Nevada winters, cougars avoided the north-facing slopes and frequented the south-facing slopes where there was less snow and more deer. When summer arrived, the cat shifted to the cooler north-facing slope, where there was more vegetation. Cougars also tended to restrict their movements to between 6,500 and 8,500 feet, where vegetation, deer, and other prey were most plentiful. (26)

Paul Beier has monitored some lions continuously for 24-hour periods to better understand their movements. He found that the lions in his California study area travel at a steady speed of approximately one-half mile per hour. Though there is a lot of variation, the cats seem to follow a cycle of traveling for a little less than an hour, followed by 30 minutes of rest. (38)

Mountain lions can cover a lot of ground in their nightly wanderings; a lion in the San Andres Mountains of New Mexico has been documented at 10 miles in a night. (17)  Not surprisingly, Hemker and his coworkers found in their Utah study area that females with kittens less than six months old moved significantly shorter distances than females with older kittens, females without kittens, transients, or resident males. One resident male traveled eight miles in a one-day period, but this represents only the straight-line distance between locations on a map and does not reflect the actual distance covered. (25)  Here is why.

mountain lion travel range

Cougars have two basic patterns of movement: hunting and traveling. When the big cat is looking for prey, it traverses a zigzag course through it home range. An entire day searching for prey likely brings the cougar only a few linear miles from its starting point. The cat is basically searching and has no specific destination in mind. A traveling cat, with a specific destination in mind is a different matter. The route will be more direct, though it will still take advantage of the landscape, following ridges, major drainages, or using low passes. (22)

A common myth about the daily movements of cougars is that they follow a regular and predictable circuit around their home range. This assumption may stem partly from observations of domestic cats or vague attempt apply human behavior to cougars. A variety of studies that monitored cougar movements with radio telemetry found no such patterns. Cougars move freely and arbitrarily about their home ranges and have never been seen to pass a fixed location at set intervals. (22)  As a stalking predator that depends on the element of surprise to capture its prey, regularity of movement would handicap the cougar’s ability to survive.

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ANATOMY OF A HUNTER Thirty-five million years of evolution have honed Felis concolor into an almost perfect predator. The cougar’s keen senses, muscular body, and remarkable adaptability make it ideally suited for a predatory existence. Because they are the most exclusive of meat-eaters, almost every feature of a cat’s body is related to the way it detects and catches its prey . (1)

DETECTING PREY The big cats use all of their senses in their ongoing search for prey. When the Florida panther stepped in front of my truck that night in the Everglades, and the eerie glint of its eyes flashed back at me, I was witness to a universal feature of feline legend. But the eyes of a cat are not windows to the fires of Hell, as witch hunters in the Middle Ages believed; (2) they are actually the marvelous adaptation of a nocturnal hunter

Cats have extraordinary vision.

Cats have extraordinary vision. The eyeball, pupil, and lens are proportionately larger than other carnivores. The eyes of a domestic cat are only slightly smaller than those of humans, but the cat can open its pupil to a maximum area three times larger than humans can; (3) this increases light-gathering ability and enhances night vision. Cougars are both nocturnal and crepuscular (active during the twilight hours of dawn and dusk). As a result, their eyes are proportionately smaller than the mostly nocturnal lynx, making pumas suited to hunting both in daylight and at night . (4, 5) The amount of light entering the eyeball is controlled by the pupil, and the pupil in smaller cats is elliptical.

After passing through the pupil and lens, light strikes a layer of light-sensitive cells at the back of the eye called the retina. There are two kinds of light receptor cells in the retinas of mammals: rods and cones. Rods function in low levels of light and do not detect color, while cones are sensitive to high levels of light and are used in color vision. Not surprisingly, the eyes of cats consist primarily of rods, though there is a concentration of cones near the center of the retina as in our own eyes. There is evidence that cats can only discern the color of relatively close or large objects, but much remains to be learned about color vision in cats . (3) Behind the retina is a thin layer of reflective cells called a tapetum lucidum , meaning “bright carpet.” The tapetum reflects light back through the rods and further enhances interpretation of low-light images, giving the cougar a second chance to discern what it is looking at. The tapetum is responsible for the characteristic “eye-shine” of reflected light that is frequently seen in mammals at night . (1, 2, 6) It was the Florida panther’s tapetum I saw that night in the Everglades.

Another characteristic of feline vision is that the eyes are close together and facing forward. This allows the field of vision of the two eyes to overlap, resulting in binocular vision. Binocular vision provides excellent depth perception and the ability to precisely judge distance; cats have the most highly developed binocular vision of all carnivores . (1) The cat’s total visual field spans 287 degrees, with a binocular overlap of 130 degrees. Humans, by contrast, have a binocular overlap of 120 degrees in a total visual field of 200 degrees. The cougar’s depth perception is most sensitive within a range of 50 to 80 feet, a critical adaptation for a predator that stalks its prey and attacks from a short distance . (6)

While a cougar’s extraordinary vision seems to give it a distinct advantage over its prey, nature has a way of compensating for advantages. In humans, the high concentration of cones in our eyes allows us to resolve (discern) visual detail in daylight, while the low concentration of rods inhibits our ability to resolve detail in low light. This resolving power is called visual acuity. While the concentration of rods and the presence of a tapetum has increased the cat’s sensitivity to low light, it has sacrificed visual acuity. Rods do not allow for much discrimination between light wavelengths, and the tapetum further blurs the image the cat sees. As a result, cats’ vision at night is six times better than that of humans, but humans have better visual acuity . (1, 2, 3)

Further, in prey species such as deer, the eyes are mounted on the side of their head. This arrangement does not allow binocular vision but does increase the total field of view and the ability to detect predators . (7) Cats also have a heightened sensitivity to movement, and biologists believe that it is the movement of prey that triggers the puma to attack. This may explain why prey typically will “freeze” after detecting a predator . (6) In the world of predator and prey, there seems to be a defense for every offense.

Although little research has been done on hearing in cougars, it is known that domestic cats can detect their prey by sound as well as sight. They can hear frequencies in the ultrasonic range and are able to move their small, rounded ears together or independently to isolate these sounds. It is also believed that an enlarged auditory bullae (the portion of the skull surrounding the middle ear) may enhance a cat’s sensitivity to certain sounds . (1, 3, 8)

Experts speculate that in most cats, vision and hearing are important for hunting, while smell plays a more active role in social behavior) In Chapter Three: Cougars at Home it was explained that pumas have a special olfactory organ in the roof of their mouth that is employed during a flehman response, presumably to determine the reproductive condition of a female . (11) Cougars also seem to share their domestic cousin’s attraction to catnip . (8, 12) Smell, a well-developed feline sense, is not primarily used in hunting. However, “We probably underestimate how much cougars use their sense of smell,” says Fred Lindzey, “I once saw a captive male picking up the scent of a deer.” Lindzey thinks cougars probably do not use their sense of smell in hunting, but can generally use it to determine whether deer are in the area . (9) Bogue and Ferrari observed a six-month-old puma kitten follow a scent trail they laid down through undergrowth with a piece of hide from a freshly killed deer . (10) Nonetheless, dogs’ sense of smell is much more highly developed. The longer muzzle of a dog boasts almost 50 square inches of olfactory cells to the cat’s 6 square inches, and 5 percent of canine brain volume is committed to its sense of smell, compared to 3 percent in cats . (1)

Cats have an acute sense of touch, particularly with the tip of their nose, toes, and paws . (8) A cat’s whiskers are specially adapted as tactile sensors; during prey capture they are extended like a net in front of the mouth so the cat can determine exactly where the prey is to accurately inflict the killing bite . (13) Barry Lopez tells of one biologist who believes that a cougar’s paws are so sensitive that when it attacks deer in pitch black it can determine the location of the head by instantly sensing the direction its hair is growing . (14) A mountain lion is the personification of power, grace, strength, speed, and agility.

CAPTURING PREY A mountain lion is the personification of power, grace, strength, speed, and agility, largely due to heavy musculature attached to a light but strong skeleton. The majority of a cougar’s body weight is muscle and sinew, with only a relatively small portion made up of bone and organs. Long, muscular legs and a flexible backbone allow strong extended strides, while its long, heavy tail provides balance on quick turns and uneven ground . (15) Horizontal leaps of 45 feet have been recorded, along with vertical leaps of 15 feet . This ability may be partly due to the fact that the cat’s rear legs are longer than its front legs. An adaption for jumping is a valuable characteristic both for attacking prey and moving through the rugged terrain most cougars inhabit . (8) Further, the anatomy of a cougar’s front limbs allows the animal to pivot sharply without losing lateral traction, (14) an important feature when grasping large prey at high speed.

The big cat is built for speed, not endurance. Writer Jim Bob Tinsley relates that, “It [the cougar] can easily outrun a pack of dogs for a few hundred yards, but its small lungs limit the distance it can cover at full stride. When out of breath, it must seek the temporary shelter of a tree or some other natural protection.” (16) Ralph Schmidt, who works with Alberta lion researchers Martin Jalkotsy and Ian Ross, believes cougars are the fastest predator in North America. “I’ve seen cougars jump out of trees and run up a slope at an unbelievable rate of speed,” says Schmidt, with a pronounced tone of respect . (17) Apparently no cougar has ever submitted to the stopwatch, but it is telling to consider that in some parts of the western United States, cougars have been known to occasionally kill pronghorn (Antilocapra americana) , the fastest land animal in North America . (16, 18)

mountain lion travel range

Cats and many other carnivores walk upright on their toes, a stance known as digitigrade, as opposed to the plantigrade stance found in humans and bears) Webbed skin and fur between the toes muffle sound as the cat walks, and while stalking or walking on snow or a muddy surface, the hindfeet can be placed almost exactly in the track of the forefeet . (20) Cougar tracks reveal four toe pads in front of a smooth, calloused, three-lobed heel pad. Adult tracks average 3 1/2 inches in width and length, (19) with the forepaws usually larger than the hindpaws. During normal walking the claws are retracted, but during quick acceleration they are extended and used for traction . (8)

Sharp claws are important for seizing and controlling prey so the cougar can deliver the killing neck bite. To keep their claws sharp, cougars have spring-like ligaments that keep the claws retracted inside fleshy sheaths and elevated above the ground most of the time. Retraction of the claws is passive, requiring no effort on the part of the cat. In use, the muscles in the forelegs contract, which in turn protract the claws, and the cougar is ready for action. As Kiltie has noted, it probably makes more sense to call them “protractile” claws . (3)

The skull is short and round, with 16 teeth in the upper jaw and 14 in the lower jaw . (6) Cougars have a powerful bite because of the reduced length of their jaws and their large jaw-closing muscles, the temporalis and masseter. Atop its skull, the cat has a bony ridge called the sagittal crest, which provides a large surface area for the attachment of the temporalis, the larger and stronger of the two muscles, and the one that lifts the jaw up and back.

The other end of the temporalis attaches to the lower jaw. At a wide gape, such an arrangement gives the temporalis a greater mechanical advantage in driving the large canine teeth through the prey’s muscle and bone. The masseter originates on the zygomatic arch, a bony arch on the side of the skull, attaches to the outside of the jaw, and lifts the jaw up and back. The masseter is more important as the jaws close at the end of the bite and when the cat uses its carnassials during feeding. Carnassials are modified molars and premolars that act as shears to cut through tough hide and tissue, which is why cats turn their heads to the side when they are biting through tissue while feeding. Cats do not chew their food, but rather use their carnassials to cut their prey up into small pieces or strips, which are swallowed whole . (13) Even the puma’s tongue is specially adapted, with sharp, horny protuberances that help remove meat from bone and also aids in grooming . (21)

The cougar seems to tolerate the bitter cold of the Canadian north or the blistering heat of the Amazon equally well. Its remarkable physique is covered with a tawny coat that is short year-round in warmer climates but grows longer and thicker during the winter in temperate regions . (22) In temperate climates, shedding occurs in the spring . (18)

mountain lion travel range

Though the cougar’s scream is a subject of some debate, the cats are capable of a surprising variety of vocalizations. While they are quite vocal during mating, it is unlikely that cougars roam about in the wild “screaming,” as depicted in films. To do so contradicts the cougar’s secretive behavior and would also be counterproductive, as it would scare away essential prey. Their repertoire includes hisses, growls, .and whistles . (23) It appears whistles may be used as a means of communication between juvenile cougars and between females and kittens; (24) additionally, captive cougars have been observed greeting familiar humans with distinct whistles . (10)

DIET Because cougars are one of the most widely ranging cats, their diet varies depending on the prey available; almost completely carnivorous, they rarely eat vegetation . (25) As noted, deer are at the top of the menu – mule deer in western North America and white-tailed deer in eastern North America . (6) Exactly how much of the lion’s diet deer comprise varies with each state and province . (25) Mule deer are the primary prey in Oregon, Alberta, and Utah, while in Florida, white-tailed deer (Odocoileus virginianus) and wild hogs (Sus scrofa) are at the top of the menu . (26) Other large prey are taken as well. Hornocker found that cougars in the Idaho Primitive Area fed primarily on mule deer and elk (Cervus canadensis) . (27) In Nevada, the ever-adaptable lion occasionally augments its diet with wild horses and desert bighorn sheep (Ovis canadensis ) . (28) Even moose (Alces alces) have been known to fall to the powerful cat in Alberta(29) and British Columbia . (30)

Opportunistic predators, cougars also feed on a variety of smaller prey, especially in times of seasonal abundance. Columbia ground squirrels are frequently the cat’s main course during the warmer summer months in Idaho, (31) while during years of peak snowshoe hare abundance in British Columbia, over a quarter of the cougar’s diet was composed of hares . (30) In the southwestern United States, peccaries (Tayassu tajacu) are taken . (6) Additional prey includes rabbits, marmots, beaver, porcupine, an assortment of birds, domestic livestock, and even carrion, as well as other carnivores, such as bobcats, coyotes, and other cougars . (18, 32)

The puma’s dependence on smaller prey is more pronounced in South America. In Chile’s Torres del Paine National Park, pumas subsist primarily on European hares (Lepus europaeus), followed by guanacos (Lama guanicoe) and domestic sheep (Yanez et al. 1986). Biologist Louise Emmons found that pumas in the jungles of Peru preyed on small rodents, oppossums, bats, and lizards. The majority of their prey were agoutis and pacas, rodents weighing 2 to 22 pounds . (34) The big cats are able to kill and eat most animals in their home range.

When the abundance of their primary prey declines, cougars have been known to switch their diet. This occurred in Big Bend National Park in western Texas, where cougars and male bobcats usually feed on deer. When the mule deer population crashed in 1980-1981, both cats were forced to switch to peccaries and lagomorphs (rabbits and hares), the next largest prey. (35)

The diet of a cougar also varies according to its sex and age. Solitary cats seem to consume more small prey than do females with young, while transients likely focus their efforts on smaller prey until they develop the skills to hunt and kill larger prey. Females must feed young for up to 18 months, so killing large animals is a more efficient way to procure a lot of fresh meat quickly. Bruce Ackerman and his colleagues concluded that, considering the food requirements of females with young, breeding populations of cougars could not exist in areas devoid of large prey like deer . (36) How often a cougar kills a deer depends on a variety of factors:

  • • Sex and reproductive status
  • • Size of the dependent young
  • • Social status
  • • Abundance of alternate prey
  • • Rate of spoilage of kill
  • • Time of year

Estimated frequencies of kill vary from 1 deer per 10 to 14 days(27) to 1 deer per 2 to 3 days . (36) As stated previously, females with dependent kittens require greater quantities of meat, quantities that increase as the kittens grow. Ackerman estimated the following kill rates for a resident female in southern Utah:

  • • Solitary resident female – 1 deer per 16 days
  • • Resident female with 3-month-old kittens – 1 deer per 9 days
  • • Resident female with three 15-month-old kittens – 1 deer per 3 days

Having evolved as an opportunistic predator, that may go without eating for days at a time, cougars exhibit fast-and-gorge feeding behavior. Captive cougars will eat 5 to 12 pounds of meat per day, more after deprivation, which is typical of this behavior . (25) Sensitive to spoilage, cougars seem to prefer their meat fresh, though they have been known to eat carrion when near starvation. In the warm temperatures of Arizona, spoilage of the carcass will typically restrict a cougar’s use of a kill to no more than four days . (38) Kills can be fed on for longer periods in more temperate regions. This is a decided advantage during cold winters when metabolic demands are high. One researcher in Idaho observed a cougar remain with an elk carcass for 19 days during one particularly cold winter . (31)

A cougar goes about its feeding with almost surgical precision.

After killing a deer, a cougar goes about its feeding with almost surgical precision. The cat plucks the fur from the point of incision using its teeth, then, using its claws, the flank is opened behind the ribs. The stomach and intestines are pulled out and dragged away from the carcass; heart, lungs, and liver are removed and eaten first . (6, 38) These organs contain higher concentrations of protein, fat, and vitamins than does muscle tissue. The liver in particular contains a high proportion of vitamins, amino acids, and glycogen, the primary source of metabolic energy. The fact these organs are consumed first is an important characteristic of the cougar’s fast-and-gorge feeding behavior. It may be a survival adaptation in case continued feeding is not possible; (39) a young transient cougar risks losing its kill to a resident lion, so the transient may gorge then leave the vicinity of the kill . (40) When continued feedings are possible, they include the rear quarters, then muscle tissue on the inside of the legs. If it’s been a long time since the last meal, the lion will feed heavily, consuming up to ten pounds of meat . (38) Generally left uneaten are the head, large bones, hide, and digestive tract. A solitary cougar can consume 75 percent of a carcass by weight, (39) while a female with kittens will perform an even more thorough job of consumption . (38)

Because they feed on more easily digestible meat than all other carnivores, cats have relatively short digestive tracts.

Both the small and large intestine are shorter and less efficient than in the more omnivorous dog. It is known, for example, that domestic cats have a mean digestive efficiency for total energy of 79 percent, compared with 89 percent for the domestic dog . (1) While the physiology of cougar digestion is poorly understood, the cat seems to survive on a diet rich in protein and fat. Veterinarian Melody Roelke reports that Florida panthers generally pass all food matter within 36 hours . (41) Houston has suggested that since the hunting behavior of cats requires quick acceleration, they minimize body weight and inertia by having a short, light gut . (42) Unlike dogs, cats appear to be unable to tolerate low levels of nitrogen (from protein) in their diets . (3)

Little is known about the water requirements of pumas. Stanley Young observed that “When water is scarce the puma apparently is capable of existing for long periods without it. This seems particularly evident in some of the dry semi-desert areas of southwestern Utah.”(12) Kitchener believes that although cats have access to free water, part of their liquid requirement probably comes from their prey . (1) Kenney Logan agrees. “There is no doubt cougars drink free water. They visit springs and guzzlers (man-made water catchments). They probably get a lot of water from drinking the blood of prey. We find very little blood spillage in the kills on our study area.” Logan believes blood provides the cougar with nutrients, vitamins, and water . (43)

PREDATORY BEHAVIOR Cougars are ambush predators . (44) Like most cats, with the notable exception of the cheetah, they attempt to catch their prey unaware, rather than chase it down. Unlike a bear, which kills its prey through brute force, the cougar is the epitome of speed and precision. The cat is silent on approach, quick on the attack, and efficient in making the kill . (15, 38) Few people have ever seen a mountain lion make a kill in the wild and a great deal of myth surrounds how it is done . (38) Stories of cougars killing 800-pound steers and scaling 10-foot fences with the unfortunate bovine still clutched in its jaws originated in frontier imaginations rather than any documented incident. Paul Leyhausen has done extensive research on predatory behavior in domestic and wild cats, including pumas . (13) It is now believed that prey-capture behavior is very similar in all species of wild cats . (1) In cougars, the process begins with the hunt.

The cougar is a relentless hunter. The search for prey is driven by the cat’s hunger and, in the case of a female, the need to feed growing kittens. The hungrier the cat, the greater the tendency to roam, with effort focused on areas where prey was previously found . (13) The cougar navigates its home range in a zigzag course, skirting open areas and taking advantage of available cover . (31, 38) The cat’s keen senses are focused to pick up the slightest movement, odor, or sound. How frequently the cat encounters prey depends on the number of prey in its home range, the density of cover, and the cougar’s searching behavior . (6) Once prey is detected, stalking begins.

The cougar fixes its gaze on the animal, lowers itself to the ground in a crouch, and begins to maneuver closer to the animal, taking care to remain hidden. It assumes an alert watching posture: head is stretched forward, the whiskers spread wide, and ears erect and turned toward the front. The cat will hold this position for minutes on end following the prey’s slightest movement with its head . (13) When the prey draws within 50 feet or less, Felis concolor strikes . (38)

Flattened against the ground, the cat darts forward, either running or with several bounds, and quickly closes the distance to its prey . (13) The angle of attack is usually from the rear or side . (1) Once at its prey’s side it grasps the neck and shoulders with the front paws, claws extended. It will frequently strike its prey with such force during the final charge that in the case of a large animal such as deer or elk, the prey will be knocked off their feet. Although cats will attack from elevated positions such as a tree or boulder, they almost never land directly on their prey. To do so would provide a very unstable landing area and a poor platform from which to counter the unpredictable movements of its prey. When attacking large prey, it is important for the puma to keep its weight on its hind legs so that it can adjust its position to maintain control, or if necessary, to make a prudent retreat to avoid injury . (13) (It has already been noted that cougars are occasionally injured and even killed during such attacks.) Instead the cat lands on the ground short of the prey and attacks from there . (13)

mountain lion travel range

Following a successful kill, cats rarely feed immediately . (13) This characteristic is probably a response to the level of hunger, energy expended during the attack, and the excitement of the attack. At some point, though, the cougar will drag or carry the carcass to a protected spot, such as under a tree, and begin to feed . (38) The cougar’s instinct to hide its kill is strong. After eating its fill, the cougar will hide the carcass by covering it with pine needles, limbs, and small twigs. Hiding the carcass protects it from scavengers, such as coyotes and ravens, and keeps the meat cool and fresh. Using its claws as rakes, the cat will stand over the kill and drag the debris inward with its front feet. Even soil and small rocks are used when nothing else is available. Shaw once found a deer killed on a large granite boulder in Arizona by a radio-collared lion. The female cougar had placed a single twig on the carcass before leaving . (38)

The cougar will typically remain in the vicinity of its kill for up to five days, making frequent trips back to feed and protect the carcass from other carnivores and scavengers. It’s not uncommon for a lion to move the carcass after each feeding. Fred Lindzey reports that lions will sometimes move a carcass over 100 yards each time, then feed on it . (9) How long the cat remains with the kill and the degree to which it is consumed depends on the size and weight of the cougar, the size of the prey, and weather conditions . (38) It has already been noted that a female with kittens will consume the carcass more completely than a solitary cat . (39) What causes abandonment of a kill is unknown. Some are fed on only once, leaving much of the meat uneaten, others are devoured completely, including the bone marrow. Spoilage and disturbance of the carcass are the most likely causes. Whatever the reason, the cat will leave the immediate area and eventually begin the search for its next prey.

In nature, predation is the rule, not the exception.

EFFECTS OF COUGAR PREDATION ON PREY In nature, predation is the rule, not the exception. All animals compete for the resources their environment provides and there are few animals that are not subject to some kind of predation . (46) Prey populations are influenced by many factors, some of the more important being the availability of food, denning sites, disease, migration, emigration, and predators . (1) The specific impact predators have on prey populations is one of the least understood and most controversial areas of study in wildlife science. Because an in-depth discussion of predator/prey relations is beyond the scope of this book, the following section gives only a brief overview of the relationship between cougars and their prey

Traditional thinking was that predators slaughtered everything in sight and were capable of decimating entire prey populations. It was believed that by eradicating the predators, the prey populations would rise, leaving more game for the human hunter. As a result, the Scottish wildcat was almost eradicated by the early 20th century, large predators in African game parks were “controlled” until recently, (1) and attempts were made to exterminate the mountain lion. The puma’s occasional habit of killing live-stock didn’t help the situation . (6)

The next evolution in predatory/prey theory suggested that predators weed out old or sick individuals, thereby improving the health of the prey population . (1) This is sometimes referred to as sanitation. While the sanitation theory has elements of truth, it is frequently a distorted and over-simplified interpretation of Darwin’s theory of natural selection . (38) Cougars are known to prey on both healthy and sick animals, (27) and whether the cat selects for unhealthy prey is generally considered unproven. Both healthy and weak are vulnerable because of the cat’s ambush-hunting method . (25) Shaw has pointed out that some aspects of the lion’s behavior suggest that they may actually tend to select for younger and more active animals – that lions are somehow triggered to attack active prey . (38) Current thinking is that cougars select prey more at random; as opportunistic hunters they will kill what they can, sick or healthy, as the opportunity arises. While circumstance and opportunity seem to be more important than conscious selection on the part of the cat, studies indicate certain segments of the prey population appear to be more vulnerable than others.

Mountain lions were found to kill a greater proportion of mule deer fawns and mature bucks during studies in Colorado (47) and Nevada . (48) Hornocker observed the same pattern in lion-killed elk in Idaho . (27) The Boulder-Escalante cougar study in southern Utah revealed that adult males (all seasons), fawns (in Winter), and older deer (7+ years) were most vulnerable to cougar predation . (37) Cougars in the Diablo Mountains of California appear to take more of a cross section of the deer population. The cats showed a preference for bucks over fawns, and did not appear to select for old and infirm . (44)

Bucks may be more vulnerable because they are more distracted during the mating season. They are in poor physical condition during the winter and have a tendency to winter at higher elevations and in broken terrain, the preferred habitat of the cougar. Unlike the more solitary bucks, females winter in loose groups, which increases their ability to detect predators. Should the group be attacked, the confusion of many fleeing animals reduces the cat’s chance of success. The very young and very old are less vigorous and thus more vulnerable as a result . (6, 25) Other factors have been noted by research biologists. Ackerman believes mule deer vulnerability is determined by their behavior, such as group alertness, parental care, and differing habitat use . (37) Susan de Treville has suggested that in the brushy chaparral habitat of California, bucks with antlers may hesitate to “bust brush” for fear of getting hung up. Consequently, they were more frequently seen in the open . (49)

The effect of predators on prey populations is not always clear. “The question becomes not ‘Do pumas limit or reduce prey growth rates?”’ explains Rick Hopkins, “but ‘When do they and under what circumstances?”’ Hopkins points out that predator/prey interactions are very complex and depend on:

  • • Prey numbers
  • • Predator numbers
  • • Prey characteristics
  • • Predator characteristics
  • • Abundance of alternative prey (50)

In the past, it was widely believed that large predators such as wolves and cougars held the numbers of deer, elk, and moose in check and that consequently, predators were critical to prevent population explosions. In the case of cougars, studies in Idaho, (27) Utah, (39) and California (44) indicate cougar populations did not significantly limit the size of the deer or elk herds. Hornocker believes the cats provide the more subtle benefits of reducing wide fluctuations in deer populations and keeping deer herds on the move by scaring and chasing them into areas that are not over-browsed. He also believes cougars cull the sick, weak, young, old, and abnormal from the deer herd and that this reduces the possibility of disease and the passing on of less-desirable genetic characteristics to future generations . (27)

Renowned biologist Paul Errington believed that removal of some prey animals by predators increased productivity in the rest of the prey population by reducing overall competition. Thus, predation compensates for initial deaths by improving the chances of survival for other members of the herd. This concept, called compensatory mortality, is generally supported by research and is frequently cited in wildlife management circles as the theoretical rationale to support recreational hunting . (51) Current evidence seems to indicate that the losses of healthy or unhealthy deer to cougars are at least partly compensatory in nature.

Science can only take us so far in our examination of the cougar. Objective and empirical methods have revealed much about the lions’ biology, but this is only one facet of the feline enigma. A broader perspective can be gained by going back in history, to a time when the division between fact and fable was less clear. It was a time before science, when the lives of Felis concolor and Homo sapiens were more intertwined than today. It was a time when the American lion was a god.

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  • Lindzey, F.G., B.B. Ackerman, D. Barnhurst, T. Becker, T.P. Hemker, S.P. Laing, C. Mecham, and W.D. Van Sickle. 1989. Boulder Escalante cougar project final report. Utah Division of Wildlife Resources, Salt Lake City, Utah.
  • Danvir, R.E and F.G. Lindzey. 1981. Feeding behavior of a captive cougar on mule deer. Encyclia. Utah Academy of Sciences. 58:50-56,
  • Roelke, ME. 1991. Veterinarian, Florida Panther Recovery Project, Florida Game and Fresh Water Fish Commission, Gainesville, Florida. (Personal communication)
  • Houston, A., C. Clark, J. McNamara, and M. Mangel. 1988. Dynamic models in behavioral and evolutionary ecology. Nature, 332:29-34. (Cited from Kitchener 1991.)
  • Cunningham, E.B. 1971. A cougar kills an elk. Canadian Field Naturalist. 85:253-254. (Cited from Dixon 1982.)
  • Whitfield, P. 1978. The hunters. Simon and Schuster, New York.
  • Dixon, KR. 1967. Mountain lion predation on big game and livestock in Colorado. Job Completion Rep. Proj. W-38-R-2l, Colorado Game, Fish, Parks Dept., Fort Collins, Colo. (Cited from Dixon 1982.)
  • Ashman, D. 1977. Mountain lion investigations. Job Performance Report, Project W-48-8. Nevada Dept. Wildl., Reno, Nevada. (Cited from Dixon 1982.)
  • Errington, P.L. 1967. Of predation and life. Iowa State University Press, Ames.

EARLY PERSPECTIVES Cougars tread a path that winds through the mythic and earthly worlds of Native American cultures throughout the Western Hemisphere. In the mythic world, the puma was the protector of the cosmos. (1)  The Zui of New Mexico said that the ancient ones wanted the world to be guarded by those keen of sight and scent, (2)  and the puma was the sentinel of the north. (3, 4, 5)  The Miwoks of California described the cougar as the ideal hunter, strong and brave, chief among the animals. (6)  The Apaches and Hualapais of Arizona regarded the cats wailing as an omen of death. (7)  In one Navajo legend, the hero is critically wounded when witch objects are shot into his body. It is the puma who extracts these missiles of death, saving the hero’s life. The Navajo also believed the puma benefited them by leaving the greater portion of their kills for the people to eat. (8)  However, not everyone viewed the cougar with reverence the Papago envisioned the lion as a flesh-eating monster. (9)

In the earthly world, many cultures believed they could imbue themselves with the pumas hunting prowess through the use of societies or fetishes. A society is a group within a tribe, usually of men, bearing a particular animal is their totem and source of power. (1)  The Caiyek (Cougar Society) of Zia Pueblo in New Mexico existed to assist hunters in the chase. Members of the Caivek believed that a supernatural ability such as successful hunting was given to each species of animal by the Powers. (10)  At Jemez Pueblo, the Cougar Society provided deer and rabbit. (11)  The cougars skill as a hunter could also be extended to warfare. (1)  Membership in the Opi, a prestigious warrior society of Chchiti Pueblo, could only be obtained by killing an enemy in battle or by killing the great hunter–the puma. (10)

In some cultures, fetishes are important in transferring supernatural powers to humans. A fetish is a material object such as a small carving, believed to contain or manifest specific power or powers. The Hopi and Zui believed that fetishes were once actual beings and Zui hunters carried small stone-carved mountain lion when they went deer hunting. The figure was fed every day, and upon the hunters return, blood of the quarry was smeared on the muzzle of the fetish. At San Felipe Pueblo, mountain lion fetishes were used before a buffalo or deer hunt. (1)

The cougar was also believed to possess great magical and medicinal power. Aztec physicians prescribed pricking the breast of sick tribesmen with the sharpened bone of a puma to ward off death. Some tribes drove away illness by Jangling dried cougar paws over a sick individuals head. Cougar gall was administered in extreme illness to increase the power of the ailing person to resist the disease, and ultimately to instill the cats ferocious fighting spirit. (7)  Certain warriors tied lion paws and muzzle hair to their bandoleers before going into war, (11)  and the Zia Cougar Society used leggings and effigies made of puma hide in certain rites. (12)

The pervasive influence of the cougar can also be found in Native American art and architecture. The ancient Peruvian city of Cuzco was laid out in the outline of a puma and skilled artisans fashioned golden puma figurines. (13)  A six-inch wooden statuette, unearthed on Marco Island, Florida, in 1895, bore the distinctive form of a living panther, and was believed to date between A.D. 1400 and A.D. 1500, when the area was occupied by the Calusa. (14)

In contrast to the reverence shown by many early cultures, Inca rulers and their subjects hunted the puma as game at the height of their civilization in ancient Peru. (7)  Even so, the cat was rarely hunted for its meat. Stalking and killing a cougar, using only a spear or bow and arrow, required beating the master hunter at its own game. Few hunters possessed such skillbear and deer no doubt proved far easier quarry. (1)

mountain lion travel range

The impact of early cultures on mountain lion populations was probably minor, even though some tribes regularly killed pumas in order to gain entrance to a warrior society or to obtain materials for ceremonial or magical use. Acuff gives three reasons for this: One, the animal was difficult to hunt. Two, the puma was revered, and one does not kill a sacred animal wantonly. Three, human population densities were low enough that such hunting had minimal impact. (1)

To native people whose survival depended on consistently procuring adequate food, the ability of the lion to kill game with apparent ease was enormously respected, and subsequent deification of the puma is not surprising. (1)  The big cat enjoyed the awe and respect of native people who shared the forests, mountains, and deserts. But in the 14th century, when the first Europeans reached the Americas, their perspective of the cougar was to be quite different.

Early explorers and settlers were driven by a desire for wealth, glory, adventure, and a tremendous religious fervor. Many of these people came from countries that were heavily settled and relatively predator-free. The Christian obsession with morality greatly influenced the Europeans view of native cultures and the wilderness that greeted their arrival. They saw it as their moral duty to civilize not only the savages but the land itself. There was no place for predators in such a world.

The cougar was not only immoral, but a competitor that vied for the abundant game of the New World and a threat to domestic livestock. During the early Spanish mission period Jesuit priests in Baja California were offering natives one bull for every cougar killed. In 1684, Connecticut offered a bounty of twenty shillings apiece for the killing of catamounts, the local name for the cougar. Massachusetts was paying bounties on cougars in 1742. (14)  Where Native Americans offered the big cat respect, the new European immigrants felt only fear and loathing. By imposing human ethics upon wild predators it was easy to make the step from viewing them as competitors to viewing them as enemies. (15)  Such simplistic thinking separated animals into two categories: beneficial (edible wild game and livestock) and injurious (all other mammals and birds).

Some native individuals did not completely acquiesce to this new vision of the sacred cougar. The Indians of peninsular California refused to kill or disturb the puma, even at the insistence of the Jesuit priests; it seems the uneaten portion of the cats kill had long been an important source of food for these people. (7)  In the early 1900s, western author Zane Grey wrote of an incident in Arizona in which where a Navajo guide refused to participate in a mountain lion hunt because that would have been tantamount to hunting a deity. (16)

From 1500 to 1900, little factual information was accumulated about the cougar. In this vacuum, the outrageous was accepted as true. Hunters and authors spun tales of a supernatural creature, more terrifying than in any of the Indian myths. Recurring themes of cowardice, gluttony, brutishness, sneakiness, and wantonness dominate. White hunters projected themselves as courageous heroes destroying an evil creature that embodied traits that were considered human vices. The white man created his own mythic wandone in which the cougar was assigned malevolent attributes in order to justify his real-world extermination. (1)

As colonization spread west, the great hardwood forests of the Northeast were cleared and subdivided, effectively destroying the cougars habitat. Alexander Crowell killed the last panther in Vermont in 1881, (17)  and the last panther in Pennsylvania was shot in 1891. (18)  By 1900 the cougar was effectively exterminated east of the Mississippi. As domestic cattle and sheep were released across the open plains of the West many of the native bison, pronghorn, elk, deer, and bighorn sheep were displaced. (15)  Because cougars found livestock increasingly available and easy to kill, it took an occasional sheep, goat, or cow. Such depredation, consistently exaggerated by stockmen, did little to enhance the cougars public image. Prevailing opinion at the turn of the century was best summed up by Theodore Roosevelt when he described the cougar during a hunt as …the big horse-killing cat, the destroyer of the deer, the lord of stealthy murder, facing his doom with a heart both craven and cruel…  (19)  Stockmen agreed, and declared war.

PREDATOR CONTROL In early 1990, several major national magazines and newspapers carried the grisly photograph of 11 severed mountain lion heads stacked under a tree. The photograph was taken anonymously by an angry Arizona state wildlife employee. The heads represented only one-fourth of the 44 lions killed in Arizona in 1989 by professional hunters working for the U.S. Department of Agricultures notorious Animal Damage Control (ADC) program. (20)

The U.S. Government entered the business of exterminating wild animals (many of them on public lands) in 1915, when western stockmen pressured Congress to appropriate $125,000 to wipe out wolves and coyotes and supposedly save beef for our allies in World War I. (21, 22)  The U.S. Biological Survey, the predecessor of the U.S. Fish and Wildlife Service, was charged with the responsibility of hiring hunters and trappers to do the job. But it was the passage of the Animal Damage Control Act of 1931 that gave birth to ADC and provided the money and authority to expand the destruction of mountain lions, wolves, coyotes, bobcats, prairie dogs, gophers, ground squirrels, jackrabbits, and other animals injurious to agriculture, horticulture, forestry, husbandry, game, or domestic animals, or that carried disease.  (21)

Between 1937 and 1970, federal employees of Animal Damage Control (ADC), derisively branded All Dead Critters by some of their critics, killed 7,255 cougars; 23,830 bears; 477,104 bobcats; 50,283 red wolves; 1,744 lobo wolves; 2,823,000 coyotes; and millions of other animals. After 1970, control was focused primarily on cougars, coyotes, and bobcats, because the grizzly bear and wolves were placed on the endangered species list. (23)

Federal predator control efforts are augmented by state hunters and bounty programs. Arizona originally considered the cougar an undesirable predator, and 2,400 were killed between 1918 and 1947. Efforts to eliminate the cat were accelerated in 1947 when the state began to offer a bounty varying from $50 to $100 per lion; between 1947 and 1969, over 5,400 cougars were slaughtered in Arizona. (24)  Federal, state, and private hunters killed 1,775 pumas in Colorado between 1916 and 1965.  (25)  California paid out bounties on 12,452 cougars killed between 1907 and 1963, when the program was eliminated by the state legislature.

A good portion of this grisly total was due to the efforts of Jay Bruce, California’s official lion-hunter from 1914 to 1942, (1)  credited with killing almost 700 cats. (18)  British Columbia lays claim to the greatest carnage, with 16,633 cats slaughtered between 1910 and 1955, (26)  The bounty on cougar in British Columbia continued from 1910 to 1957; during that time the total kill probably exceeded 20,000 animals. (27)  According to statistics compiled by Ronald

Nowak of the U.S. Fish and Wildlife Service, a minimum of 66,665 cougars were killed within states and provinces between 1907 and 1978. (26)

The most common methods used to control cougars were poison, trapping, and hunting with dogs. (28)  At the turn of the century, predator control consisted of guarding livestock using people or dogs, as well as leghold traps, snares, and guns. Poisons were not introduced into wide use until the l930s. The predator control arsenal contained such deadly elements as thallium sulfate, strychnine, and cyanide. The notorious poison, sodium monoflouracetate, commonly known as Compound 1080, appeared in the late 194Os. It was injected into bait carcasses, which were then transported to bait stations by vehicle or dropped from the air. Poisoned bait stations killed coyotes, bears, and eagles by the thousands. In 1972, President Nixon issued an executive order banning the use of poisons to kill predators on public lands, and the Environmental Protection Agency (EPA) disallowed the registration of such toxicants. Presidents Ford and Carter agreed with the ban, but President Reagan subsequently rescinded the original executive order, and EPA once again began registering 1080 for use in the early 1980s. (23)  Today, 1080 toxic collars, which are fastened around the necks of sheep, are used to prevent attacks in Texas, Montana, Wyoming, New Mexico, and Idaho. ADC also uses strychnine for underground rodent control. (29)

Due to the cougars low density and preference for killing its own prey, poisons were generally ineffective in eradicating it. Poisoning of lion-killed carcasses proved the most effective technique, though the kills are difficult to find. Young transient cats are probably most susceptible to poisons because of their tendency to eat unusual prey or foods. (28)

Steel-jawed, leghold traps and snares are still used today in cougar control programs though, like poisons, they are of limited effect. The cats do not respond well to scented bait traps, so hunters use what are called blind sets: unscented and camouflaged traps set in canyon bottoms and along rims in areas cougars are known to travel. Foothold snares can also be baited or unbaited. Some trappers even use catnip with some success. The traps or snares are then checked on a regular basis and, if a cat is found in one, it is shot. The biggest limitations to traps and snares are the time, expense, and labor necessary to make and maintain adequate sets. (28)  Both poisoning and trapping are nonselective: many animals not specifically targeted by control efforts such as hawks, eagles, badgers, foxes, deer, Javelina, livestock, and domestic dogs have died in traps and snares set for other predators.

Today, hunting cougars with dogs is the most frequently used capture method, both by predator control agents and sport hunters. In 1988, the majority of cougars killed by ADC agents in California, Nevada, Oregon, and Utah were tracked down with hunting dogs. (30)  Specially trained hounds are used to track, then chase the big cats. Though a dog is no match for a cougar in speed or combat, the cats seem to consistently retreat from barking dogs. If tracking conditions are good, such as after a light snowfall, and the dogs pick up a fresh scent, the cat is no match for the hounds endurance and will quickly seek refuge in a tree. While in the false safety of the tree, the cougar is then shot.

Until 1986, ADC was a branch of the U.S. Fish and Wildlife Service. Following the recommendation of a 1970 presidential commission that the predator policies should be eliminated or reduced on public lands, (21)  and, due to a growing conservation ethic during the 1970s and 1980s, the role of the ADC was greatly reduced. Finally, western members of Congress, under the prodding of disgruntled stockmen, succeeded in getting ADC transferred to the U.S. Department of Agriculture and the program rebounded, The toll in 1989 included 237 cougars, 236 black bears, 80 gray wolves, 1,220 bobcats, 7,158 foxes, and 86,502 coyotes, 10,000 more than in 1988. (22)

Too frequently, the fiscal logic of ADC defies reason, and the ADC program is being increasingly criticized because of its enormous expense and lack of effectiveness. As Harley Shaw states, … information available on lion population biology suggests strongly that control of lions is unlikely to occur and [is] definitely not cost efficient. (28)  In 1990 the ADC program spent $29.4 million in federal dollars$3.8 million more than 1989plus about $15 million in state funds to destroy vast numbers of mammals and birds   considered predators or pests. In 1988, California spent $3.2 million to kill 32,368 mammals about $100 for each animal. The amount of damage the animals caused to livestock, poultry, and crops was placed at $1.4 million. It cost twice as much to destroy the predators (41 cougars among them) than if the ranchers and farmers had been compensated for their losses. (20)

DEPREDATION The killing of livestock (depredation) still fuels the political fires that advocate the killing of cougars through predator-control programs and hunting. Stockmen, biologists, and conservationists consistently cross swords over one question: How much impact do cougars have on livestock operations?

The majority of depredation permits issued to kill mountain lions are for attacks on sheep and cattle, Taking California as an example, the National Agricultural Statistics Service reports that in 1988 there were 4,600,000 cattle and calves (beef and dairy) in California and 800,000 sheep (sheep and lambs). During the same year the California Department of Fish and Game issued only 102 depredation permits for attacks on cattle, calves, sheep, and lambs. (31)  In other states the situation is similar. Wain Evans states that, Verified depredations affect less than 1 percent of New Mexico ranchers each year. (32)  Suminski reports that in Nevada, the estimated annual losses of sheep to cougars averaged 0.29 percent. (33)  Balancing these numbers, Fred Lindzey emphasizes that although depredation is a small problem industry-wide, local ranchers can be significantly affected. (34)

Depredation statistics indicate that cougars cause fewer livestock deaths than do other species, such as domestic dogs. Between June 1986 and June 1987, ADC information for 41 California counties showed that cougars were responsible for 5 percent of all sheep and lambs lost to predators, while domestic dogs caused 11 percent of sheep deaths and coyotes 78 percent of deaths. (35)  In southwestern Utah, coyotes caused 92 percent of sheep losses and cougars were responsible for 7 percent. (36)

While to humans, cattle are inappropriate prey for a mountain lion, to the opportunistic hunter, killing available and vulnerable prey, whether deer or cattle, is how the cat stays alive. In that context killing livestock is perfectly normal cougar behavior. Cougars will kill most species of domestic livestock, though cattle and sheep are common prey. Arizona and New Mexico suffer the most frequent losses of cattle, with incidences generally decreasing northward through the cats range. This pattern may be largely due to ranching practices, with cattle and calf losses greatest in areas where they are born in cougar habitat. (38)  However, Susan de Treville points out that cattle and lions were plentiful throughout her study area in the Big Sur region of California, yet there was little depredation, except for a few goats killed by an elderly male lion. (39)  Lindzey also emphasizes that no one has demonstrated a relationship between lion density and depredation. (34)  Sheep losses occur anywhere they are grazed in cougar habitat, but are greatest in the summer when the sheep are dispersed on open range. During the winter, sheep are moved to lower ground and often to fenced-in pastures. (40, 41)  Hopkins points out that if mountain lions were completely neutral in the prey they select, livestock would be killed in proportion to how frequently they occur in the cats home range. Evidence shows that cattle and sheep are infrequent prey, indicating lions generally avoid them. (42)

Most cattle killed by cougars are calves less than one year old and weigh less than 200 pounds. Though they have been known to kill cows up to 800 pounds, experts feel kills in excess of 300-400 pounds are unusual. (41)

Sheep of all age classes are killed by cougars, although lambs seem to be taken more often if they are present. (36)  Cougars will kill more than one sheep in a single incident, though it will only feed on one or two. A lion in Nevada once killed 59 sheep (mostly lambs) in one attack. (33)  Such occurrences of surplus killing are easily interpreted as evidence that the cougar is a bloodthirsty predator that enjoys killing. This may be true: to enjoy killing is probably necessary to survive. But like so much of the cats behavior, it is not likely to be that simple.

Why does surplus killing occur? It is partly because the drive to kill an animal outweighs the need to eat it, (43)  and because domestic livestock cannot or will not escape. Experiments have shown that both hungry and satiated felines continued to kill prey as long as it was presented rather than eat those already killed, (43)  To a cougar, multiple kills are an efficient way to procure a lot of food in a short period of time, and are wasteful only in an artificial captive situation where prey animals are kept at very high densities and are unable to escape. (44)  Guggisberg explains that the cats urge to pounce upon a victim is constantly being reactivated by the penned-in animals helplessly milling around it. The situation it finds itself in is quite abnormal, and so, of course, is the pumas reaction. It would never be able to perform a massacre of this kind among the wild animals which form its natural prey, for they take flight the moment one of a herd has been struck down. (45)

Traditionally, there have been two approaches to cougar depredation: kill the problem lion or reduce the lion population in areas where attacks occur. (41)  Most states (including California) and provinces have laws that enable livestock owners to protect their animals. Depredating lions can be destroyed if caught in the act of attacking livestock, or they can be captured and killed if the depredation incident can be verified.

Cougar population reduction is usually attempted through sport hunting hunters are directed to specific areas experiencing livestock losses. Such general population reductions are usually ineffective. Harley Shaw advises that in good lion habitat, attempts to control lions at any feasible level will probably accomplish nothing. (28)  The complete elimination of cougars from problem regions or zones has been tried three times in New Mexico twice to protect domestic sheep and once to protect wild sheep. None of these removals resulted in a reduction of depredation. (32)  In areas where grazing is seasonal, sport hunters generally remove cougars in the winter, but livestock are not moved to these areas until summer. This provides ample time for new transient cougars to move into the area and take up residence in the hunter-created vacuum, resulting in a cougar population equal to or greater than the original population. (41)  If killing resident cougars in a population results in an increase of transients, and transients are more prone to take unusual prey than resident cats, it is likely that aggressively hunting the cats could actually increase depredation.

Nonlethal predator control methods are available that reduce depredation. One of the most effective ways is to change livestock management practices. Economic conditions have led some ranchers to shift from sheep operations to cattle operations. In areas experiencing heavy losses to cougars, cattle ranchers have changed from cow-calf to steer operations. (36)  Lindzey advises, Pasturing livestock, primarily sheep, in more open areas and avoiding timbered areas particularly those in steep, broken terrain should reduce livestock losses to cougars. (46)  For small farms raising sheep and goats, electric fences have proven effective against predators, including coyotes. (47, 48)  Keeping sheep more tightly herded reduces predation, but it also increases range deterioration. (36)  Some sheep operations have experimented with guard dogs; these breeds are different from herding dogs and include large types such as Great Pyrenees and the Hungarian Komondor. Dogs raised with sheep and properly trained offer continual livestock protection that adapts to changes in predator behavior. (48, 49)  However, guard dogs have proven less effective where sheep herds are scattered thinly over the range. (36)

mountain lion travel range

Two states, Colorado and Wyoming, pay ranchers compensation for cougar depredation. The lion was classified as a game animal in Colorado in 1965 and the Division of Wildlife (DOW) became liable for damages at that time. Although the original legislation was contested, it was upheld by the courts; (36)  Colorado paid $44,959 in compensation for livestock losses between 1990 and 1991, according to Cathy Moser of the DOWs Terrestrial Section.

John Talbott, Assistant Chief Game Warden for the Wyoming Game and Fish Department (WGFD), explains that his department is responsible for damage to domestic livestock by trophy game animals, and that the cougar became a game animal in 1981. Most of Wyomings losses are sheep rather than cattle, and not surprisingly, the biggest point of contention between state wildlife officials and ranchers is the magnitude of such losses. WGFD rarely pays for the entire loss. No compensation is allowed in either state if a landowner does not permit hunting on his property.

Talbott believes Wyoming’s compensation program increases tolerance of the cougars and helps ranchers view them as a bona fide wildlife resource that people like to see out there. Both he and Moser think the programs increase cooperation between property owners and the states, Talbort further points out that the expense of compensation programs is not just in the money paid to ranchers. There are also the substantial costs of monitoring both sheep and lions, verifying losses, negotiating compensation, and administrative time. On the negative side of the issue is the question of whether a public agency managing a public resource, frequently on public land, should be reimbursing ranchers for losses of private property.

HUNTING The 1960s brought a new perspective on predators and a limited degree of protection for the cougar as state legislatures and wildlife managers in most of the western states shifted the big cats official status from injurious predator to game animal. Nevada reclassified the cougar as a game animal in 1965; Washington did so in 1966, followed by Utah in 1967 and California in 1969. All other western states followed suit. Two Canadian provinces, Alberta and British Columbia, did likewise. California set a new precedent in 1990 when residents passed a referendum giving the cougar complete protection from sport hunting (Proposition 117). At the other extreme is Texas, where the lion is still considered a varmint and receives no protection from the state whatsoever. (50)

Hunting seasons vary in length from year-round to one month, with bag limits usually 1 cougar per season per person. Most states require a permit. Northern states usually set their seasons in late fall and winter, when snow provides the best conditions for tracking with dogs. Some states time their hunting seasons so they do not conflict with other big-game seasons and to reduce conflicts between cougar hunters and their dogs and people hunting deer and elk. Other states time seasons to avoid cougar birth peaks. This is thought to provide a measure of protection for small, vulnerable kittens and presumably reduces the number of young kittens orphaned when their mothers are killed (more on this later), Texas and Oregon also allow trapping as a method of sport harvest. (41)

A number of states offer special pursuit-only seasons. During pursuit-only seasons, hunters are allowed to chase and tree cats, but not to kill them. This provides a way for hunters to train and condition their dogs and is also used by wildlife photographers to get a picture of this elusive animal in the wild; however, some experts point out that cougars can still be injured or killed during such seasons. Lindzey explains: Permanent physical damage to a mountain lion may result from prolonged or frequent chases. Also, small kittens are as vulnerable to being killed by dogs during these seasons as they are during regular sport-hunting seasons. (41)

Successful cougar hunting requires the use of specially trained tracking hounds and sometimes involves several days spent trailing the elusive cats through miles of rugged terrain. Since few hunters are willing to make such an investment in time or effort to train dogs, they usually employ the services of a hunting guide. For a fee of $1,500 to $5,000, the guide supplies the food, lodging, hounds, horses, four-wheel-drive truck, and familiarity with the local terrain. Some hunting guides use CB radios, walkie talkies, and fit their hounds with special radio collars, which allow the guide and hunter to radio track the hounds while the hounds trail the cougar. The radio collars frequently contain a switch that is triggered when the dog tips its head back for an extended period, indicating that a cougar has been treed.

While most hunting guides run legitimate operations, the expense of long pursuits and the impatience of clients to bag a trophy cat entice some guides to provide a higher level of convenience in the form of will-call (as in, When we have your cougar treed, we will call you) or shootout (as in, All you have to do is shoot it out of the tree) hunts. (50)  The guide puts a list of clients in his pocket, then heads out into the woods to find and track a cougar. Once he has a cat treed he radios the client or leaves the cat under the watchful eye of a helper and drives to the nearest telephone. The client then flies and drives to the location of the treed cougar to collect his trophy. As a result, cougars remain up in the tree for days at a time, under a death watch. If it jumps from the tree it is simply treed again, until the client arrives. (Bears are also frequent victims of will-call hunts.) Although keeping a cougar in a tree overnight does violate some state laws that prohibit hunting at night, this form of hunting is not illegal. Many question the morality of these will-call hunts, though. Bill Powers, coordinator of Arizona’s Operation Game Thief silent-witness program, says Id guess that around 30 percent of the guided hunts in Arizona are will-call hunts, and that’s probably conservative. (50)

Cougar hunting is growing in popularity. In those states that allow lion hunting, the number of permits has risen steadily over the last decade. Permit sales have more than doubled in Utah during the past ten years, with little or no increase in the number of lions killed. (46)  Utah charges state residents only $12 for a license, plus a $27 lion permit fee, Nonresidents must pay $40 for the license, and $252 for a lion permit. A mountain lion hunting license in Colorado costs $30 for a resident and $250 for a nonresident. Revenues from license and tag sales rarely cover the costs of cougar research and management programs. Most of the money to be made from cougar hunting goes to guides, with local economies probably receiving a portion of the revenue. (41)

With so much time, energy, and money expended on cougar hunting, one question eventually presents itself: What impact does hunting have on mountain lion populations? Its a simple enough question, but asking it sparks immediate debate between hunters, anti-hunters, and non-hunters. Unfortunately, such debates generate more heat than light, and when these factions turn to wildlife biologists, no simple answers are available. Surprisingly little effort has been devoted to gaining an understanding of how sport hunting affects cougar populations. (46)  (For updated information, see MLFs 2006 study on the affects of cougar sport hunting in MLFs Library.)

Many hunters, wildlife managers, and biologists insist the hunting of cougars necessary to keep cougar populations healthy, prevent overpopulation, and protect deer herds. None of these arguments is based on sound biology. The primary justification recreational hunting in contemporary society is based on notion that wildlife populations produce a harvestable surplus of individuals, (51)  and that removal of this surplus benefits the population as a whole by reducing competition. For example, by killing a portion of a mule deer herd there is less competition for forage within the remaining herd, resulting in less starvation. Thus, the initial deaths due hunting compensate for, or reduce other forms of natural mortality such as starvation. Biologists such as Paul Errington believe each species population has surplus individuals whose death is assured, regardless of the literal cause of death. Whether the surplus individuals die by starvation, predation, or hunting makes no difference, and advocates of this compensatory-mortality theory view hunting as a substitute form of predation. (52)

While most recreational hunting programs are based on this theory of harvesting the surplus, and a significant body of research supports it, many of the studies were conducted on species such as deer, muskrats, and quail, animals that exist in large numbers and exhibit large population fluctuations. It may be more difficult to apply such reasoning to pumas, low-density predators whose populations appear to fluctuate very little and whose population dynamics are poorly understood.

Hunting cougars probably never completely compensates for natural mortality, and in some cases may even add to it. Lindzey and his colleagues found this to be the case while studying survival rates of mountain lions in southern Utah. Hunting mortality will not be fully  compensated for  [emphasis added] by a reduction in other sources of mortality. Intraspecific killings [fighting] and debilitating injuries resulting from attempts to capture prey will occur even in populations where densities are reduced by hunting. The degree to which hunting is  additive  [emphasis added] to other forms of mortality needs to be identified…. (53)

Sport hunting advocates point to the evidence that cougar populations appear to tolerate tremendously high levels of hunting mortality and still recover. Kerry Murphy monitored the response of a small population of western Montana lions to heavy sport hunting. He concluded that over 50 percent of the resident adults in the area were killed, but were replaced by young adults raised in the area or transients immigrating from surrounding areas. (54)  David Ashman and his coworkers in Nevada believe that even under removal of 30 to 50 percent of lion populations, the cats are capable of rapidly replacing annual losses. (55)  While these examples certainly testify to the pumas resilience, they tell little about the effect of hunting on the cats intricate population dynamics.

Another study in southern Utah indicates that, when hunting mortality is combined with high natural mortality, cougars may not be so resilient. Biologists simulated a cougar hunt by removing 27 percent of the cats from a population, Seven lions were placed in captivity or transported out of the study area the first year, while another three died of natural causes the following year. By the second year the population had failed to recover to its pre-removal level. The investigators concluded that Failure of the population to recover to pre-removal level two years later and its inability to replace one of three resident adults that died the second year, suggests the population would not have recovered as quickly from a second years harvest of similar intensity. The effect of hunting on cougar populations will depend both on level of harvest and sex and age of cougars removed; populations will be most sensitive to loss of resident adult females. (56)

John Laundre and Donald Streubal, who are studying cougar ecology and behavior in Idaho, wonder about the effect of hunting on kitten mortality. The fact that males kill kittens, [has been] documented in earlier studies as well as ours. According to evolutionary theory, males would not kill their own kittens, writes Streubel. (57)  Therefore, if hunting selects males from the population, new males would be more frequently moving into new home [ranges] and thus increasing chances of non-relatedness with resident kittens. This would seem to increase the mortality of kittens.

Most states prohibit killing females with kittens or spotted kittens. However, it may be difficult for hunters to differentiate females with kittens from other mountain lions because young are often not with their mothers. Biologist Dan Barnhurst observed that the tracks of newborn to 6-month-old kittens were found with their mothers only 19 percent of the time, while tracks of 7-to-12-month old kittens were found with their mothers tracks 43 percent of the time. Tracks are the sign most used by hunters, but even if a hunter wished to comply with the laws and avoid killing spotted kittens or females with kittens, 80 percent of the time he would have no way of knowing that the female has kittens. (58)

Barnhurst also examined the vulnerability of cougars to hunting. For kittens less than six months old, being orphaned is probably the main source of mortality in heavily hunted populations. A second source of hunting-related kitten mortality is that of kittens being killed by dogs. The risk of this is greatest for kittens less than three months old, because the mother must return frequently to the den to nurse them, The tracks she leaves will eventually lead dogs to the kittens, Since the kittens are too young to climb trees or outrun the dogs, they may be caught on the ground and killed before the hunter is aware of their presence. (58)  This situation emphasizes why even pursuit-only seasons can result in lions being killed.

mountain lion travel range

Unfortunately, the current sport hunting season in many states begins in the fall and it is  probable that in some areas, cub mortality from maulings and orphaning is as significant as adult harvest  [emphasis added], writes Thomas Hemker and his colleagues. The degree that hunting and non-hunting deaths compensate for each other is not clear, although it seems that causes of natural mortality are at least partially independent of hunting. Consequently, it seems probable that cub survival would be lower in hunted populations where either sex of adult cougars may be removed. (59)

Experts hope that some better answers to these perplexing hunting questions will be provided by a 1O-year research study currently underway in the San Andres Mountains of southern New Mexico. Because the mountains lie within the U.S. Army’s White Sands Missile Range, cougar hunting has been prohibited. It will be the first long-term intensive study of un-hunted cougars in a desert environment. Biologists Kenney Logan, Linda Sweanor, and Frank Smith of the Wildlife Research Institute in Moscow, Idaho, are manipulating the resident lion population by removing a portion of the cats to simulate an overharvest. After studying the general ecology of the lion population for five years, they removed 70 percent of the lions (13 adults and sub-adults) from the southern one-third of the San Andres Range. The cats were then translocated to an area in northern New Mexico, where another biologist will monitor their activity. Logan says the project has four objectives:

  • To determine the effects of the removal on lion population dynamics. This includes the immigration of transients, emigration (dispersal) of kittens, changes in the behavior of the lions toward each other, and general distribution and movement of the lions in the study area.
  • To determine the effects of the removal on lion social organization. This includes changes in the ratio of males to females, changes in the age classes of lions (residents, transients, and kittens), or shifts in home range size and overlap.
  • To determine the effects of lion removal on the survival of desert mule deer ( Odocoileus hemionus crookeri ) and desert bighorn sheep ( Ovis canadensis mexicana ), which the lions prey on in the San Andres.
  • To monitor the behavior of the translocated mountain lions. Scheduled for completion in 1995, the project is a cooperative effort between the New Mexico Department of Fish Game, the U.S. Fish and Wildlife Service, and the Sands Missile Range. (60)  (For updated information, see a summary of the Logan/Sweanor study in MLFs Library)

Existing research provides little evidence that sport hunting benefits cougar populations by keeping them healthy. As to whether lions overpopulate or devastate herds, studies indicate that the cats social behavior generally keeps their numbers in check (28, 61)  and that their impact on healthy deer populations is thought to be minimal. Nor does hunting mountain lions reduce incidents of depredation. (36)  Mountain lion sport hunting exists today primarily to provide a recreational experience for hunters want to bag a trophy.

One fact is certain, however: 2,176 cougars died at the hands of hunters in western North America in 1990. Idaho lead the pack with 350 lions killed, followed by Alberta 254, and Montana at 228. Sport hunting is still the greatest known cause of death in cougars.  (For more information see MLFs reports on mountain lion mortalities in MLFs Library.)

POACHING November 16, 1990, state and federal authorities converged on a ranch near Lockwood, California. What they were the skulls, heads, and hides of mountain lions, tigers, spotted leopards, black leopards, and jaguars remnants of illegal hunts conducted by the owners, Floyd and Dawn Patterson. (63)  Big game hunters paid the Pattersons up to $3,500 for the privilege shooting the big cats and taking their stuffed carcasses as trophies. Most of the animals were thought to been surplus zoo animals and many were simply shot in the stock trailer they were delivered in. One cat was dragged out with a lasso around its neck and shot just outside the door. (64)  The Pattersons were tried and convicted on 42 counts of violating state wildlife laws. (65)  Federal agents are still investigating the crimes and may yet file additional charges, which carry even more severe penalties.

The effect of poaching on cougar populations in North America is unknown, but according to the U.S. Fish and Wildlife Service (FWS), trade in illegal animal parts is a $200 million a year business, and growing, a 100 percent increase since 1980. John Turner, FWS Director, says, Our wildlife is under siege. (66)  The international trade in wildlife, both legal and illegal, is a $5 billion-a-year business, and the United States is the largest consumer, accounting for $1 billion of the revenue annually. (67)

Illegally trapped cougars and endangered jaguars are purchased by unscrupulous outfitters for hunters willing to pay big trophy fees. (68)  Cougars and bighorns are killed for their hides and heads. Bear gall bladders are sold as aphrodisiacs and their jaws, teeth and claws for jewelry. Elk and deer are killed and their hides, meat, and antlers sold. Major purchasers of poached goods are Asian medicine shops and markets. In the Orient, animal parts such as antlers and bear and cougar gall bladders are believed to have strong curative powers.

No longer simply an occasional deer taken out of season or a couple of fish over the limit, the age of large-scale commercial poaching has arrived. Skilled, organized, and well-equipped teams of poachers are decimating our nations wildlife and reaping obscene profits in the process. According to Larry Farnsworth, commercial poaching is a nationwide problem of a considerable magnitude. The data … indicates that commercial poaching is one of the most underreported crimes in the country. Farnsworth estimated the minimum value of the illegal sale of wildlife at $175,100,773 and feels this number is extremely low. (69)

National parks, once seen as refuges, are becoming the killing fields of our wildlife. Obsessed trophy hunters in search of the biggest prizes are slipping into parks to shoot mountain lions, elk, deer, grizzlies, and bighorn sheep. Their reward is an entry in a record book, wall mounts, pictures in albums, and quick profits. (68)  Black bears are slaughtered by the hundreds in Great Smoky Mountains National Park; hunters in Alaska’s Denali National Park use small airplanes to gun down entire packs of wolves for their valuable pelts; poachers in Rocky Mountain National Park target bighorn sheep, deer, and elk. Rangers in Alaska consider illegal trophy hunting their worst problem. (67)

Poaching is considered by some experts to be less of a threat to cougars due to the difficulty of hunting lions without dogs and the low value of the pelt; however, this may be changing. In Los Angeles, a Korean buyer placed orders for 300 bear paws, 14 bear gall bladders, and 15 cougar gall bladders. (70)  The California Department of Fish and Game reports there were 43 known cougars illegally killed between 1984 and 1990. In 1988, the Washington Department of Game states that of the 121 cougars killed in the state, 32 of them were illegal. (30)  With bear gall bladders going for $540 per ounce and a record-sized bighorn sheep head worth $50,000, the incentive to poach is high. It is also important to note that cougar gall bladders are visually indistinguishable from bear gall bladders.

Heavy poaching of deer herds can have indirect impacts on cougars by removing potential prey. Based on information provided by California, Oregon, Idaho, Michigan, Nevada, Texas, Washington, and Wisconsin, roughly one-half or more of some fish and game species that are taken each year might be taken illegally by poachers. California Department of Fish and Game estimates, for example, that of the approximately 160,000 deer taken in California in 1984, roughly 100,000, or 62 percent were killed illegally. (71)

Poachers face little risk of being caught. There are only 7,200 state game wardens and fewer than 200 federal wildlife agents nationwide, to enforce the myriad wildlife laws. (72)  According to Pete Bontadelli, former director of California Department of Fish and Game, California has 330 game wardens in the field, which comes out to about 1 warden for every 300,000 acres. A past study showed we would need five times the current warden force to do the job effectively. (73)  Bontadelli says their poaching apprehension rate is about 4 percent, which is higher than some states that estimate their violator apprehension rate between 1 and 2 percent. Most violations are not even reported. Canadian wildlife officials hired a man in 1987 to commit a variety of hunting violations, including poaching. Of the 762 crimes, only 8 (1.1 percent) were ever reported. (67)

Poachers also face little chance of severe punishment should they be caught. While Floyd Patterson was sentenced to six months in jail and fined $28,200 and his wife Dawn was fined $14,100 and sentenced to 200 hours of community service, (74)  they were the exceptions. Most courts, straining under an overwhelming docket of crimes such as murder, rape, robbery, and drug-related crimes tend to look upon wildlife violations as minor infractions. Guilty violators usually walk away with small fines or a suspended sentence. Under such circumstances, poaching has the ominous potential to become yet another obstacle to the cougars survival.

COUGAR ATTACKS ON HUMANS American folklore is filled with stories of the Devil Cat descending from the mountains, announcing its presence with a bloodcurdling scream, and searching for any opportunity to drag helpless victims off into the night. Such stories have all the elements of good fiction. Unfortunately, the shy and secretive nature of the cougar helped to nurture these fables. By the early 1900s a different version of the mountain lion was slowly beginning to emerge. Tracy Storer wrote in 1923 that, It is the general belief among naturalists and well informed laymen that the … Mountain Lion … does not ordinarily attack human beings. This belief is strengthened by the experience of many thousands of people who have camped and lived in the range of the mountain lion without being harmed in any way. (75)  Early puma researcher Frank Hibben claimed, Man, who upon greater acquaintance with the cougar loses much of his dread of the animal … [will] find a curious, gentle, and very likeable disposition supplanting the vicious side. (76)  Even Teddy Roosevelt changed his view, claiming that it would be no more dangerous to sleep in woods populated with mountain lions than if they were ordinary house cats …  (77)

There is no question that cougars attack people on occasion. The earliest recorded fatal attack in the United States is thought to have been a 58-year-old man living in Chester County, Pennsylvania, in 1751. (7)  The most recent was an 18-year-old jogger killed near Idaho Springs, Colorado, in January 1991. (78)  In order to assess the risk lions pose, other important questions must be answered: How frequently do these attacks occur? Why do they occur? What can be done to prevent future attacks?

In an attempt to get some answers, Paul Beier conducted a detailed review and analysis of cougar attacks on humans in the United States and Canada. Only those cases where a cougar bit, clawed, or knocked down a human were considered. Maulings by captive cougars and cases in which a person (e.g., a cougar hunter) deliberately approached or harassed a wild cougar were excluded. Each report was included only if it could be verified by a newspaper or other published account that included statements from medical personnel, law enforcement officers, wildlife officials, or park rangers. Applying this criteria, Beier found that from 1890 to 1990, there were 53 recorded cougar attacks in the United States and Canada. There were 9 fatal attacks resulting in 10 deaths and 44 nonfatal attacks resulting in 48 injuries. The greater number of victims is because there were 2 victims in 5 of the attacks. (79)

Children were found to be more vulnerable than adults, making up 64 percent of the victims. Those 5 to 9 years old were in the highest risk group. Whether or not the child was with an adult also seemed to influence their vulnerability. Of 37 child victims, 35 percent were alone, 43 percent were in groups of children, and 22 percent were accompanied by adults. Adults made up 36 percent of the victims, and the majority (11 of 17), were alone when attacked. (79)

Underweight yearling lions (12 to 23 months old) seem more inclined to attack people, and made up 40 percent of the offending cats. At this age an immature cougar increasingly hunts without help from its mother and by 14 to 24 months of age it moves into a new and often unfamiliar home range. (55, 61)  Under these stresses, some yearlings may have difficulty capturing wild prey, writes Beier. The body mass of most yearling attackers suggests that this may be a factor. Two of the underweight yearling attackers had porcupine quills in their throat. In addition, the report shows that half of attacking adult lions were noticeably underweight.

Only two offending cougars were found to have a disease or injury. One was probably rabid and caused two deaths from a single attack near Morgan Hill, California, in 1909. This is the only known case of rabies being transmitted from a cougar to humans; both victims died of the disease, not from the physical injuries. (75)  The second diseased cougar had cataracts and was killed near Cowichan Lake, British Columbia, in 1916.

British Columbia accounts for over half (57 percent) all recorded attacks, with the hot spot being Vancouver Island, a 12,408-square-mile island with 300,000 human residents. Twenty of the 53 attacks (38 percent) took place on Vancouver Island, 10 on mainland British Columbia, 5 in Texas, 4 in California, 3 each in Alberta and Colorado, 2 each in Arizona, Montana, and Washington, and 1 each in New Mexico and Nevada.  (79)  Experts are at a loss to explain why there is such a concentration of attacks on Vancouver Island. Dennis Pemble, a British Columbia wildlife-control officer, thinks the reason is because of the islands lush and heavy cover, which allows the cats to stalk close without being detected. (80)  Others have observed that smaller prey, such as porcupines, cottontail rabbits, and opossums are absent from the island. This lack of small prey may prove stressful to yearling lions less proficient at taking deer, and may contribute to attacks on humans. (79)

Across the Georgia Strait, on mainland British Columbia, Pemble is frequently called upon to kill or capture and move mountain lions that have wandered into the suburban neighborhoods of North and West Vancouver, a city of almost a half-million people. Excellent deer and cougar habitat exists only a few miles from the center of the city, and occasionally lions will take a pet dog or cat. Pemble notes that most of the cats he captures in or near Vancouver are young animals or adult males. (81)  In July 1991, two children and their teacher were mauled by a lion near the town of Lillooet, about 120 miles northeast of Vancouver. (80)  Seidensticker notes that what impresses him is not the number of attacks in the Vancouver area but how really few there have been, given the close proximity of mountain lions and people. (81)

Beier’s study further points out that aggressive behavior on the part of intended victims may discourage a lion about to attack or even repel an attack in progress. This differs from the more passive practice of playing dead or curling up in a fetal position during bear attacks. Because the cougar is such a skilled stalker, few victims actually saw the cat before being clawed or bitten. In those, individuals who did have time to react, shouting, swinging a stick, waving arms above the head, or throwing rocks clearly deterred the cougar from carrying out an attack. It also seems advisable not to run. In at least two cases, running appeared to stimulate the cougar to chase and attack the victim. The majority of people with whom pumas actually made physical contact fought back with bare hands, a stick, a knife, a jacket, or a rock. These efforts usually succeeded in driving off the cat. Even children who were alone were able to repel the cougar by fighting back. (79)

In a recent issue of Smithsonian, Seidensticker and Lumpkin speculated as to what would motivate a cougar to attack a human. Like other big cats, mountain lions specialize in killing large mammals with hooves, primarily deer and elk. A human standing up is just not the right shape for a cats prey. An erect persons head and neck are in the wrong place, and most adults are taller than even the largest of the cougars prey species. The position of the neck is most important, for that is where the cat must deliver the killing bite, However, a person bending over, squatting, or running may present a more attractive prey configuration to a cougar. The authors point out that tigers sometimes kill Asian rubber tappers and grass cutters who bend over frequently while working, or people who go out at night and squat to relieve themselves. They suggest that perhaps the Colorado jogger who was recently killed had attracted the cats attention as he ran along, then stopped and crouched down to tie a loose shoelace. (81)

Ironically, the animal that causes more human deaths than any other is not the cougar, but the cougars primary prey: deer.

Ironically, the animal that causes more human deaths than any other is not the cougar, but the cougars primary prey: deer. According to the National Highway Safety Administration, deer were responsible for 130 human fatalities in 1989 alone; most of these were due to collisions between the animals and motor vehicles. (82)  (The number of deer killed in these accidents will be discussed in the next section.) But deer are not the only deadly threats to humans. Each year in the United States, domestic dogs kill 18 to 20 people and inflict 200,000 injuries requiring stitches, (83)  bee stings are fatal to over 40 individuals, and rattlesnakes inflict 5,000 bites resulting in 12 human deaths. (84)  Yet none of these very real hazards seem to stir near the interest of cougar sightings. Writers Dan Bensimhom and Mary Brophy suggest that it would be wise to forget about lions and tigers and bears. Just keep a sharp eye out for Bambi, bees, and mans best friend. (82)

Mountain lion attacks do seem to have increased in the last 20 years. There were more fatal attacks during the last 21 years (6) than during the previous 80 years (4). Authorities generally believe there are two reasons for the increase: mountain lion populations are growing in some areas because states and provinces have changed the cats status from bountied predator to a game species and, in the case of California, have given the puma complete protection. Simultaneously, human populations are growing, along with their use of wildlands, which has increased the potential for encounters. (79)

Shaw observes that there has likely been a marked change in the way people react to seeing a cougar in the wild. Early hunters and ranchers would not have made an official report of a sighting, or even of a threatening incident. The lion was usually shot whether it threatened or not. Modern, unarmed hikers are more likely to report a sighting. When lions show up near more populated areas the threat becomes an official problem, usually with media involvement. Shaw suspects that to some extent, were dealing with an increase in noise, not numbers. (85)

Some people have strongly suggested that attacks can be prevented through regular hunting of cougars; however, there is no evidence that cougars are more likely to attack humans in un-hunted areas. (79)  As previously mentioned, 57 percent of all attacks took place in British Columbia, yet hunters and predator control agents kill almost 200 cougars in the province annually. (27)  Additionally, the number of attacks in Texas (5) and in Colorado (4), both of which allow hunting, does not differ significantly from California (4), where sport hunting has been illegal since 1972.

In an effort to understand what is behind these increasing encounters between cougars and people, the first Mountain Lion-Human Interaction Symposium and Workshop was held in Denver, Colorado, in April 1991. Hundreds of cougar researchers, wildlife managers, conservationists (including this author), and reporters from across North America gathered to hear the presentation of papers, to participate in discussions, to compare notes, and to talk. One point frequently made was about the increasing urban/wilderness interface. Simply put, this is the growing fringe of urban areas that is pushing deeper into prime mountain lion habitat. Jim Halfpenny, a lion researcher from the University of Colorado, emphasized the need for studies on lions near urban areas. Dennis Pemble from British Columbia and several wildlife officers from Colorado Department of Wildlife spoke at length about having to increasingly respond to reported lions in suburban areas. By the end of the three-day conference it was clear that not much was known about lion behavior and encounters; however, there seemed to be agreement on one point: there is an important need to educate the public about cougars and how to live with them.

Recent events in southern California may portend what could happen if wildlife and land managers do not take an aggressive approach to educating the public about mountain lions. In 1986, two children were mauled by cougars within a few months of each other in Ronald W. Caspers Wilderness Park, east of San Juan Capistrano. The attacks were the first in California since 1909. In August 1991, the parents of the first child won a lawsuit against Orange County, which has jurisdiction over the park, and were awarded $2 million. (86)  While the victims lawyer claimed the county had been negligent in warning visitors about lions in the park, the case has ominous implications for resource agencies being held liable for the behavior of wildlife. Orange County plans to appeal the case, but the second victim is waiting in the wings with another lawsuit. Sadly, in August 1992, the Orange County Board of Supervisors banned all children under the age of 18 from Caspers Wilderness Park. (87)

HABITAT LOSS AND FRAGMENTATION When our ancestors first set foot in New England they beheld magnificent, primeval forests that would have rivaled today’s few remaining old-growth forests in the Northwest. Huge, old trees, stately white pines 200 feet or more in height, hemlocks, maples, beeches, and birches thrived among a richly diverse community of trees and shrubs, Trees of different heights and an extraordinary mix of species created an equally varied and rich range of opportunities for birds, mammals, and all other fauna. Through this paradise trod the catamount, or eastern panther, hunting the abundant white-tailed deer. (88)

mountain lion travel range

Within a mere 200 to 300 years it was all gone the forests and the wildlife. Colonial land clearing and burning for settlements was only the beginning. By the middle of the nineteenth century, most forests had been cut over two or three times, exploited for lumber, pulp, fuel, charcoal, potash, tannin, pitch, and ship stores. Market hunting, along with ongoing, unregulated hunting and trapping, resulted in the astonishing scarcity of many game species (notably deer) by the late 1700s. All predators and any animal with a marketable hide were threatened or locally exterminated by 1900. The panther, eastern timber wolf, wolverine, pine martin, fisher, and even the beaver disappeared completely. (88)

The cougar survives in western North America primarily because of 700 million acres set aside as public land. Maurice Hornocker acknowledges the importance of this habitat and credits the Wilderness Act of 1964 as an example of legislation beneficial to cougars. While the national forests, national parks, wildlife refuges, and other state and federal lands provide a measure of protection, loss of critical habitat is still the greatest single threat to wildlife and the cougar is no exception.

Urban, residential, and agricultural development encroaches on cougar habitat throughout North America. In Colorado, the growing urban corridor that extends along the eastern shoulder of the Rocky Mountains, from Boulder south to Denver and Colorado Springs, presses in on the margins of cougar country. Rampant growth on Florida’s coasts is squeezing the Everglades on both sides and threatening the few remaining panthers. The urban sprawl of Tucson and other communities is adjacent to some of Arizona’s densest cougar populations, which are located in the southeast corner of the state. Since 1945, California has lost over 5 million acres of wildlife habitat and, from 1948 to 1990, the states human population rose from 9.6 million to 30 million. New residents have a penchant for settling in the brushy chaparral country of the western Sierra Nevada and in the coastal mountains. Both areas are prime mountain lion habitat. San Diego, Los Angeles, and the San Francisco Bay Area all have adjacent mountain lion populations.

As the stress of urban living begins to take its toll, more and more people seek to escape, finding a temporary refuge in the public lands. Many of the growing cities in the West provide easy access to nearby national parks, national forests, and other public lands, and annual visitation to Americas national parks now exceeds 260 million a year. Increasing numbers of visitors backpack in the most remote wilderness areas that are home to the cougar.

The U.S. Forest Service, Bureau of Land Management, and U.S. Fish and Wildlife Service administer the public lands under a policy of multiple use. This allows such activities as logging, mining, livestock grazing, and recreational uses such as hunting, off-road vehicles, and snowmobiles. Increased logging and mining operations mean more roads, which further subdivides valuable wildlife habitat. Cougar research in southern Utah and northern Arizona showed that the cats tend to avoid areas of high road densities and recent clear cuts. (89)  Extensive dirt road networks in an area also make cougars more vulnerable to hunting. Dan Barnhurst believes that road closures would be very effective in limiting the number of cougars killed during a hunting season and could be used for specific goals such as discouraging hunting in a drainage known to be occupied by a female and kittens. He further states, Since increased road access increases cougar vulnerability, the potential impact on the cougar population should be considered in the environmental impact statement of any planned projects that include construction of new roads (i.e., timber sales and fossil fuels or mineral exploration). (58)

Paved roads are probably the most efficient wildlife slaughtering mechanisms ever devised, Each year, millions of wild animals are killed on Americas highways. The California Department of Transportation estimates that automobiles kill almost 20,000 mule deer each year, a number equal to deer killed by hunters annually. (90)  The toll these collisions have on human life was mentioned in the previous section. When combined with development, highways pose a triple threat to wildlife: as development reduces the amount of available habitat and squeezes remaining wildlife into smaller and more isolated pockets, high-speed traffic on larger and wider highways kills more and more of the remaining population. (91)  Imagine the risk a mountain lion faces in attempting to cross an eight-lane interstate freeway. Of the 32 cougars Paul Beier has collared in the Santa Ana Mountains of southern California, nine have been hit by cars. Only two survived. (92)

In the long run, these habitat fragmenting forces may be more degrading to North Americas wildlife population than actual loss of habitat acreage, says Larry Harris, a professor at the University of Florida. According to Harris, habitat fragmentation results in four major consequences for wildlife:

  •  Loss of area-sensitive species, animals whose existence and successful reproduction depend on the size of the habitat area in which they occur. The cougar, bobcat, and black bear fall into this category.
  • Large species that are highly mobile and occur at low densities under the best of conditions are lost. Again, the cougar is representative.
  • When coupled with the loss of large native carnivores, fragmented and human-altered landscapes (providing artificial sources of food and shelter) become dominated by exotic or already common species.
  • Inbreeding begins to occur in isolated populations of low density, such as the endangered Florida panther. (91)

The cougar is an area-sensitive species, requiring a home range so great as to demand large areas for sustenance. Without a sufficiently large territory it cannot find sufficient food or freely interbreed. Lack of available mates leads to inbreeding, which leads to losses of libido, fertility, and reproduction. (93)

Preservation of large tracts of natural land certainly seems to be the solution. But how big is big enough? Even our largest national parks are losing species. William Newmark of the University of Michigan has surveyed the history of local extinctions of mammals in national parks of western North America and has made a startling discovery. Since 14 western parks were established, Newmark documented 44 local extinctions among carnivores, ungulates, hares, and rabbits, the most commonly documented species. The main problem appears to be that even our largest parks are too small. (94)

Working with the isolated population of cougars in Santa Ana Mountains, Paul Beier has estimated minimum habitat areas for cougars using a computer model. According to his findings, if a population is confined to a small area and no immigration of transient lions is possible, the habitat area must be 620 – 1,370 square miles in size for the population to survive 100 years. Beier cautions that the minimum size is based on an optimistic estimate of lion density. For isolated populations to survive longer than 100 years would require constant monitoring and possible translocation of lions from other populations. For areas smaller than 620 square miles, extinction of the population is almost inevitable. However, if as few as 1 to 4 animals per decade could immigrate into a small population, the probability of the population surviving improves greatly. Thus a corridor for immigration would benefit a small population in an area where further loss of habitat will occur. (95)

Tying isolated tracts of habitat together with movement or conservation corridors is a frequently proposed solution to the rampant habitat fragmentation currently taking place. Larry Harris, an expert on conservation corridors, states that Our numerous, large wildlife sanctuaries must be made to function as a system, rather than being thought of as islands unto themselves. Physical interconnections of habitat must be developed and safeguarded if the wide-ranging mammals are to survive in perpetuity. In short, we need a system of wildlife conservation corridors to interconnect the many and sometimes large refuges already established. (93)  New Mexico lion researcher Linda Sweanor emphasizes the importance of wildlife corridors to cougars: It is apparent that fragmented habitat may only support lions on a long-term basis if individuals are allowed to successfully immigrate, hence requiring dispersal from other local populations. To conserve populations of mountain lions over the long term, adequate habitats must be maintained in an effective patchwork composed of relatively large blocks of wildland reserves interconnected by dispersal corridors. (96)

Beier’s research has shown that not only are movement corridors important to the cats survival, but the cougar is an ideal species for identification of corridor locations. This is because the cougar is an area-sensitive species dependent on the size of its habitat for existence and successful reproduction. Thus, a movement corridor identified on the basis of cougar use is likely to benefit at least one species. Secondly, a hunting cougar travels an average of 5 1/2 miles per night and as a result generates lots of corridor information in a short time. Collection of comparable information for less wide-ranging species may take years or generations. (95)

Though our past record of wildlife habitat protection is poor, some recent events give cause for hope. First of all, in June 1990, the people of California passed the Wildlife Protection Act of 1990 (Proposition 117). Proposition 117 prohibits trophy hunting of mountain lions, tightens regulations on protection of livestock from lions, and requires the state to spend $30 million annually for the next 30 years to protect wildlife habitat in California, In all, 57,761 acres of wildlife habitat were acquired in the first year under the provisions of this new law. We are proud to have helped in writing and passing Proposition 117, said Mark J. Palmer, Conservation Director of the Mountain Lion Foundation. Many exciting natural areas have now been permanently protected by the state for all people to enjoy. Endangered species, mountain lions, deer, wetlands, and riparian (streamside) zones have all benefited from this landmark new law. (97)

Officials in northern Florida are attempting to purchase tracts of land in an effort to create conservation corridors. The recent protective designation of Pinhook Swamp creates a strategic link between the much larger Osceola National Forest and Okeefenokee National Wildlife Refuge. Now the area encompasses sufficient prime habitat that it could be considered as a possible site for the reintroduction of the Florida panther. (91)

In Los Angeles, efforts are underway to protect a wildlife corridor connecting the isolated Santa Monica Mountains with the inland wilderness in Angeles and Los Padres national forests. The corridor, a narrowing swath of land, acts as a crucial habitat link for animals to replenish populations in the urban parklands. Even here, adjacent to the second-largest urban area in the United States, experts estimate a small population of less than 10 cougars survive. (98)

Finally, the northeastern woods are coming back. Forest ecologist Susan Morse, who makes her home in Vermont, explains what is happening: Today, we are literally watching the land rebuild itself and much of our former wildlife is returning, some with our help of course. This is true in varying degrees throughout the Northeast, but it is especially pronounced in northern New England where human population pressures have not been felt fur many decades. Now you can see beaver ponds strung like pearls between clean, flowing streams. Moose, deer, bear, bobcat, river otter, fisher, and fox have returned, along with the coyote, a newcomer to this region. [Some believe the eastern panther itself is back.] New plant growth is slowly repairing what was a ravaged land a mere 80 years ago. Its our second chance at paradise. (88)

Unfortunately, the comeback of the northeastern forests is facing a new threat. Over 85 percent of the roughly 26 million acres of forest land are privately held, mostly by a dozen companies, families, and trusts. Population growth, second-home development, massive clear cutting, and pressures from Wall Street on longtime owners to sell are raising the specter of subdividing the wilderness. Increasingly, the land has less value as timberland than as house lots. In the current depressed economy neither environmental groups nor the federal government has taken any concrete steps towards preservation. (99)  This too is unfortunate for, as Susan Morse points out, It will never get any cheaper or easier to take care of habitat and wildlife than now. (88) Without protective action, this second paradise could become paradise lost again.

Cougars are secretive, solitary, and highly mobile carnivores.

RESEARCH As has been discussed, cougars are secretive, solitary, and highly mobile carnivores that occur in low densities and roam enormous tracts of wilderness. For decades, these characteristics have made the species extremely difficult to research. The only sources of information about mountain lion populations were observations made and the carcasses collected by professional hunters. Jay Bruce, California’s official lion hunter from 1914 to 1942, was one of the first to record anatomical measurements of lions and estimate their home range and population sizes. Not surprisingly, his observations always showed the need for a state-supported lion hunter. (1)

Frank C. Hibben was probably the first biologist to enter the cougars domain and attempt a scientific examination. He spent a year (19341935) accompanying lion hunters throughout New Mexico and Arizona and analyzed the stomach contents of the cougars they killed. Hibbens findings showed deer were the preferred prey and that livestock appeared in only 2 percent of the lions examined. He also considered attaching metal tags to the ears of some lions to learn something of their movements, but soon abandoned the idea. [I]t was feared that this would raise too much opposition with cattle and game interests. (76)

In 1946, Stanley P. Young and Edward A. Goldman, biologists with U.S. Fish and Wildlife Service, wrote The Puma: Mysterious American Cat, the most comprehensive summary of knowledge on cougars up to that time. The book was based primarily on the records of federal trappers working in the West. (7)  During the next 15 years, researcher W. Leslie Robinette continued to examine the cougars food habits in Nevada and Utah, again relying primarily on the stomach contents of lions killed by federal and private hunters. He also wrote about cougar productivity and life history. (100, 101)

The first structured field study of mountain lions was conducted by Maurice Hornocker in the Idaho Primitive Area (now the Frank Church River of No Return Wilderness) in the mid-1960s. Working with veteran lion hunter Wilbur Wiles, Hornocker tracked and treed cougars with dogs, then sedated them with newly developed drugs administered through darts fired from an air rifle. They tracked, treed, weighed, measured, photographed, released, and recaptured dozens of the big cats through several bitter-cold Idaho winters. Hornocker’s work was the first ever to use marked mountain lions to document territoriality in any cat species. He was first to recognize and describe the role of transient individuals, as opposed to full-time residents in a population, and the ways in which such transients may recolonize adjacent areas. Further, his analysis of the effect of cougar predation on elk and deer populations is regarded as classic. (62, 102)

In the early 1970s, Hornocker and Wiles were joined by researchers John Seidensticker and John Messick. Several of the cougars in the study area were fitted with collars so their movements could be monitored. The radio telemetry work provided important information on the social organization of the lions in the Idaho Primitive Area. It also contributed to population estimates, sex, age, reproductive status, and population health data. (61)

Additional studies followed as other researchers adopted Hornocker and Seidensticker’s methods and soon more pieces of the puzzle began to fall into place. Cougar research has been conducted in every western state as well as Florida, Alberta, and British Columbia. Harley Shaws work in Arizona contributed valuable information on livestock depredation and the tracking of lions. Fred Lindzey coordinated the team of biologists on the 10-year Boulder-Escalante Cougar Project in southern Utah, which provided important data on the cats food habits, habitat use, and response to hunting. Canadian researchers Martin Jalkotsy and Ian Ross are finishing up another 10-year study in Alberta, and Paul Beier’s work in southern California may help save a threatened population of pumas from rampant habitat loss. In Florida, researchers such as David Maehr, Bass, and Chris Belden are fighting to save the endangered panther.

Today, as founder and director of the Wildlife Research Institute, Hornocker and his hand-picked teams of biologists are engaged in a 10-year study of cougars and their prey in the desert environment of New Mexico and a comprehensive ecological study of cougars in Yellowstone National Park. (103)  (Both of these studies were mentioned previously.) Hornocker’s almost 30 years of experience studying different cougar populations in a variety of habitats provides valuable insight into the variability of lion behavior.

Though many questions have been answered, many questions remain. Information is lacking in a large portion of the cougars range, particularly in Mexico and Central and South America. Wildlife managers and researchers still search for better ways to census and estimate population sizes, better sexing and aging techniques, ways to estimate population growth, ways to determine the impact of hunting on populations, and nonlethal ways to reduce livestock depredation. In many cases, basic survival information such as the size of populations, the size of habitats, and the number of kittens that survive to adulthood is lacking.

In 1988, Lynne Trulio, an ecologist formerly with the Mountain Lion Foundation, conducted a survey of 30 mountain lion experts throughout the United States and Canada, gathering the opinions of experts on research methods and perhaps helping guide scientists as they design their studies. Trulio concluded: Respondents listed the three primary problems with research today as funding, study length/intensity, and methodology problems. These problems all impede the understanding of mountain lions and prevent us from addressing the threats which face this species. (104)

Allen Anderson, in his comprehensive review of literature on the mountain lion, made the following recommendations regarding research needs: Aside from the diet and certain variables of reproductive biology, it is not possible to generalize on any aspect of puma biology. Thus, research is needed on all aspects. However, I believe research having a direct bearing on the welfare of puma should receive first priority. Since sport hunting is the major and only controllable source of recorded puma mortality, I suggest research on the effects of sport hunting on specific puma populations should receive that priority. Second priority should be concerned with … census and aging techniques…. (105)

Few research studies have monitored cougar populations over periods longer than a few years, and experts are reluctant to apply findings to cougar populations in different areas, given the cats recognized adaptability and subsequent behavioral variation. Lack of standardized research methods make it difficult for researchers to share and apply information. As a result, wildlife administrators too frequently face the task of designing cougar management programs based on research data that is both scant and questionable.

MANAGEMENT It is Aldo Leopold’s 1933 book,  Game Management , that is generally recognized as the birth of the science of game management, (106)  though many of its professionals acknowledge that the discipline is still so young that it is more an art than a science. Early game-management philosophy was similar to that of agriculture, and in many ways is still practiced today. Killing of game is a harvest and a population of animals a crop. The most common approach in most agencies responsible for game management is a technique called sustained-yield harvesting. Ordinarily, this involves killing a surplus portion of a population at a rate that balances the productivity in that population. (40)  Since almost all game managers depend on hunting license purchases as their primary source of revenue, the focus of management is usually on maximizing populations for the benefit of hunters. As a result, the partnership between the hunting community and game managers has a long tradition, and the influence of the hunting community has on game management practices is substantial, In many cases game populations are not managed by hunting, as is so often claimed; rather, populations are managed for hunting. Under such circumstances, politics frequently play a bigger role in management decisions than does biology.

Today, the cougar is trapped in the crossfire of conflicting management goals. Management goals may vary from the reestablishment of viable populations in their former range, as found in the Florida Panther Recovery Plan, to the complexity of maintaining viable cougar populations while providing hunting opportunities and reducing livestock losses (41) in Idaho, to complete protection of the cougar in California. While the wildlife management responsibility of most agencies ends at the state line, lions frequently cross them. A protected cougar in California that crosses into Nevada becomes a game animal. Yet, there is little coordination in management or research between states or provinces. Agencies even maintain their records differently. (For instance, collecting and reconciling the sport-hunting statistics for this book was a long and arduous task.)

There is little evidence that sport hunting is of any benefit to cougar populations.

There is little evidence that sport hunting is of any benefit to cougar populations. Nor does hunting protect deer herds, decrease depredation, or reduce the risk of attacks on people. Under such circumstances hunting advocates find themselves in the position of trying to prove sport hunting does not harm lion populations. The biological jury is still out on this question, but considering our past record in predator management it seems wise to err to the side of caution. A mountain lions hide is of little value and its meat is rarely eaten. The primary justification for hunting mountain lions is because hunters want to collect a trophy.

However, in a national survey conducted by the U.S. Fish and Wildlife Service, 80 percent of the American people disapproved of trophy hunting. (107)  Former California Governor George Deukmejian, the California Fish and Game Commission, and the Department of Fish and Game (DFG) all underestimated this change in attitude toward wildlife, particularly mountain lions. In 1985, Deukmejian vetoed a hunting moratorium on mountain lions that had been in place since 1972. The commission and DFG subsequently recommended a sport hunt. A coalition of conservation groups were able to stop the hunt for five years through court action. But the most dramatic public statement came on June 6, 1990, when the people of California approved ballot initiative Proposition 117 which permanently banned mountain lion hunting in the state.

Increasing environmental awareness, concern over loss of habitat, and evolving attitudes toward animals has a new generation of wildlife biologists advocating a shift from game management to wildlife management. A sense of profound change pervades the wildlife management field today, writes Stephen R. Kellert. (108)  Various indicators suggest that basic shifts have occurred in American attitudes and recreational uses of wildlife. These changes have been reflected, for example, in a series of studies of American attitudes, knowledge, and behaviors toward wildlife (107, 109, 110)  as well as in the findings of the 1980 National hunting, fishing, and wildlife-related recreation survey, (111) which estimated that a remarkable $40 billion [is] spent on all forms of wildlife recreation, including $14.8 billion on non-consumptive wildlife use. Non-consumptive wildlife use includes such activities as birdwatching, visits to zoos or museums, scientific study, or photography. (107)

mountain lion travel range

Many state wildlife agencies are being called upon to expand their responsibilities beyond traditional game management into such areas as nongame management, nature education, and environmental protection and mitigation. There are those who feel wildlife agencies should be spending more time managing the 90 percent of animal species that are not hunted, as well as acquiring and protecting critical habitat for nongame and game. Unfortunately, these new demands come at a time of tight budgets and declining revenues from hunting and fishing licenses, revenues that provide a large portion of the operating capital for many state wildlife agencies.

Nowhere is this dilemma better represented than in California. A recent study of the California Department of Fish and Game (DFG) by the Legislative Analysts Office (112) made the following observation: DFG historically has provided services and programs primarily for those that use or consume the states wildlife and natural habitat resource, such as individuals who hunt and fish. As California’s population has grown, leading to increasing urbanization, this traditional constituency group of the DFG has diminished steadily. Meanwhile, the responsibilities of the DFG relating to general habitat protection and endangered species protection have increased, requiring the DFG to expand services and programs that protect the overall resource base. What is lacking, however, is a clear focus on exactly what the DFGs relative priorities are…. The report makes a number of recommendations to the State Legislature to resolve ongoing problems within DFG. At the top of the list is: Determine the primary mission of the department so that conflicts between programs focused on resource use and programs focused on resource protection can be resolved. (112)

Our current wildlife habitat crisis makes it imperative that we also shift our management emphasis from species to spaces.

Our current wildlife habitat crisis makes it imperative that we also shift our management emphasis from species to spaces. For instance, few cougar management programs are coordinated with deer management, yet deer are the primary food source for the cats and suffer the same consequences of habitat loss. The future of both species is irrevocably linked, while at the same time their interactions are poorly understood. The extensive data being assembled by scientists about mule deer could prove valuable in cougar management as well. Detailed maps of the summer and winter ranges of migratory deer herds could provide a good template for preliminary identification of cougar habitat. Inventories of preferred forage for deer and the location of kills made by cougars could add to the limited information available on the favored stalking cover of the big cats. Such integrated management would be beneficial to the many other plant and animal species found within deer and mountain lion habitat.

In a 1983 report to the New Mexico State Legislature, Wain Evans, assistant director of the State Game and Fish Department, recommended that the best approach to cougar management was to allow populations to stabilize and to reimburse ranchers for the few losses to cats. Evans states in his report that The cougars biology render their populations uniquely unsusceptible to most forms of wildlife management. Efforts to reduce depredations on livestock and wildlife through cougar hunting and control on problem areas have failed. While of biological origin, depredations of livestock are essentially a political concern. Cougar management should recognize that present know-how and technologies are not sufficient to artificially calibrate cougar populations short of extirpation. Management should take advantage of the cougars self-limiting potential by allowing development of stable social structures over most of the occupied range … a system of reimbursing ranchers for at least part of their loss should be developed. (32)

It is time wildlife managers look to the cougar as more than a revenue-generating game animal or a pest that occasionally preys on livestock. Kellert reminds us that if the wildlife profession is to avoid increasing isolation from the millions of Americans primarily interested in non-game wildlife, dramatic changes in traditional programs will be required, (108)  As the wildlife managers constituency expands and diversifies, he can expect management decisions to be subject to even greater scrutiny. Fred Lindzey points out that, It is easier to quantify the dollar value of livestock losses than the recreational and esthetics values associated with mountain lions. He warns that management programs that provide for sport hunting and the killing of cougars to protect livestock will be exposed to increasing scrutiny in the future, and that Management agencies must be prepared to document that such removals will not result in a loss of the population. (41)

A PERSONAL PERSPECTIVE Finally, coexistence with cougars is possible, but it requires changing our attitude toward the wild animals that share our landscape. Living in cougar country poses some risk, but its a manageable one. Part of Florida’s success in bringing the alligator back from the brink of extinction was an aggressive public education campaign about the importance of making room for wildlife. Both the Colorado Division of Wildlife and the Montana Department of Fish, Wildlife and Parks have produced excellent brochures on how to live with mountain lions, and the Mountain Lion Foundation works hard to get the facts out about cougars. Renowned puma biologist Stanley Young once said, The almost universal fear of the puma is based mainly on its mysterious ways, size, and power to do harm, not on its aggressiveness, for as a rule it is notoriously timid in relation to man. (7)  Perhaps its time for man to give a little.

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  • ____. 1961. Notes on cougar productivity and life history.  Journal of Mammalogy  42:204-217.
  • Hornocker, M.G. 1969a. Winter territoriality in mountain lions.  Journal of Wildlife Management  33:457-464.
  • ____. 1992. Founder and Director, Wildlife Research Institute, Inc., Moscow, Idaho. (Personal communication)
  • Trulio, L. 1989b. What mountain lion scientists say about their research: Results of a survey on mountain lion research methods. Mountain Lion Preservation Foundation, Sacramento, California.
  • Leopold, A. 1933.  Game management . The University of Wisconsin Press, Madison, Wisconsin.
  • Kellert, S.R. 1980.  Activities of the American public relating to animals . U.S. Government Printing Office #024-010-00-624-2, Washington, D.C.
  • ____. 1985. Birdwatching in American Society.  Leisure Sciences  7(3):343-360.
  • ____. 1979.  Public attitudes toward critical wildlife and natural habitat issues . U.S. Government Printing Office #024-010-00-623-4, Washington, D.C. (Cited from Kellert 1985.)
  • ____ and J.K. Berry. 1981.  Knowledge, affection and basic attitudes toward animals in American society . U.S. Government Printing Office #024-010-00-625-1, Washington, D.C. (Cited from Kellert 1985.)
  • U.S. Department of the Interior, 1982.  1980 National Hunting, Fishing and Wildlife-Related Recreation Survey . Washington, D.C. (Cited from Kellert 1985.)
  • Hill, E.G. 1991.  A review of the Department of Fish and Game: Issues and options for improving its performance . Legislative Analyst’s Office. Sacramento, California. 3 September, 1991.

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A Tale of Two Big Cats: What We Learned About the Mountain Lions in Illinois in 2022

Sightings of mountain lions ( Puma concolor ) are rare now in Illinois, but historically they were one of the top predators of deer and other game in the prairie state. Mountain lions were eliminated from the state before 1870 due to habitat loss and unregulated hunting. While there have been recent confirmed sightings, currently there is no evidence of resident breeding populations of mountain lion in Illinois. Since 2002, there have been eight confirmed occurrences of mountain lions in Illinois. This includes two male mountain lions that separately made their way to Illinois in 2022. And as their populations increase in South Dakota, Nebraska, and the Rocky Mountain states, it is likely that individual mountain lions will continue to disperse through Illinois looking for new territories.

Habitat suitability map for mountain lions in Illinois. Researchers predict that only seven percent of Illinois lands could support a mountain lion.

Will these large predators find suitable habitat when they reach Illinois? A study conducted at the Cooperative Wildlife Research Laboratory at Southern Illinois University (SIU) found that less than seven percent of land cover in Illinois provides suitable habitat for mountain lions, and none of that is contiguous habitat.

That second point is noteworthy, since large expanses of suitable habitat are more likely to support breeding populations of large carnivores. The SIU researchers concluded that Illinois can provide “stopover habitat” for large carnivores such as mountain lions, gray wolves, and black bears, but that expanses of suitable habitat to support breeding populations are not plentiful.

Of course, mountain lions dispersing to new areas don’t know what lies ahead of them in their search for potential mates and good hunting grounds. Here is what biologists know about the two mountain lions that entered Illinois in 2022.

On September 28, 2022, a mountain lion was captured on a trail camera on private property in Whiteside County and the photo evidence was later confirmed in the field by wildlife biologists. Then on October 16, 2022, a mountain lion was struck and killed by a vehicle on Interstate 88 in DeKalb County. Given the proximity of the two locations, just 70 miles apart, it is thought that it was likely the same animal. Although it cannot be confirmed, this mountain lion may also have been the same animal that Wisconsin Department of Natural Resources biologists had been monitoring moving south through Wisconsin during the summer and fall of 2022.

At the time, the Illinois Department of Natural Resources (IDNR) reported that the mountain lion was transferred by the Illinois State Police to an IDNR Wildlife Biologist and then delivered to the University of Illinois Champaign-Urbana (UIUC) for a full necropsy. The UIUC necropsy results found that the mountain lion was a male in poor condition. Its stomach was empty, and it had pneumonia as well as parasites.

DNA was also collected and submitted to the U.S. Department of Agriculture (USDA) U.S. Forest Service Rocky Mountain Research Station for analysis. The genetic analysis was recently completed and provides valuable information to biologists about the animal, its place of origin, and insight into the exploratory movements of mountain lions across the Midwest. Results of the DNA analysis show that this mountain lion was a new individual that had not previously been documented or represented in the mountain lion genetic reference database. The USDA lab conducted assignment tests, which are probabilities that an animal is from a particular population based on gene frequencies and indicates how it relates to other sampled individuals from a particular area or population.

Field camera image of a mountain lion at night. The animal is to the right in the image with its eyes shining. It is walking throuh tall grass with a shrub in the background.

The genotype from this mountain lion was determined to have a low likelihood of being from any of the sampled reference populations. The USDA lab database includes over 3,000 samples from mountain lion populations in South Dakota, North Dakota, Nebraska, Idaho, Montana, Wyoming, Colorado, New Mexico, Arizona, Texas, Oregon, and Florida. The lab does not have samples from other areas with known mountain lion populations, including Washington, California, Nevada, Utah, or Mexico and Canada. Since this mountain lion did not match any of the known populations in the database, it could have been from one of the states or countries not represented in the database. Another possibility is that the mountain lion was from one of the states with samples in the database but came from a previously unsampled population. Mountain lions travel great distances and breed throughout their range, so gene frequencies throughout a particular state are likely similar. For now, it seems that this mountain lion’s origin story will remain a mystery.

Unlike the first mountain lion, the origin of the second mountain lion that moved through Illinois last fall is known. In October 2022, the Nebraska Game and Parks Commission (NGPC) informed the IDNR that a mountain lion that NGPC was tracking was within the city limits of Springfield, Illinois. The NGPC had attached a GPS collar to that mountain lion in November 2021 as part of an ongoing research project on their mountain lion population. USDA Wildlife Biologists and IDNR Wildlife Biologists and Conservation Police Officers monitored the mountain lion’s movement patterns for two days as it moved through a wooded area on the edge of the city. On the third morning, the mountain lion moved into a densely populated portion of the city and was found to be in a small backyard surrounded by homes, an apartment complex, and busy streets.

mountain lion travel range

A tough decision had to be made. The mission of the IDNR is “To manage, conserve and protect Illinois’ natural, recreational and cultural resources, further the public’s understanding and appreciation of those resources, and promote the education, science and public safety of Illinois’ natural resources for present and future generations.” The question facing the biologists was: What is the best way to manage and protect a large carnivore that is on the move while also educating people and protecting public safety? There is not a simple, one-size-fits-all answer to such a question.

In this case, it was determined that, given its proximity to an urban area, the mountain lion may have posed a potential threat to people, and the difficult decision was made to capture and relocate the mountain lion. The mountain lion was immobilized by staff from the U.S. Department of Agriculture’s Wildlife Services and sent to the Exotic Feline Rescue Center in Center Point, Indiana. The male, known as NE 110 or the Nebraska/Illinois couga r, received veterinary care and was found to be in good condition.

The tale does not end there of course. Obviously, managing large carnivores like mountain lions is more complicated than managing other species of wildlife. Knowing when and where large carnivores travel across the state will allow wildlife managers to proactively address potential human-carnivore conflicts. To assist with that, the IDNR requests that the public report mountain lion sightings in Illinois.

A mountain lion facing the camera. A large tan cat with white whiskers and greenish eyes.

It is crucial that sightings are reported as soon as possible. Mountain lions can travel long distances in a short amount of time, and weather events can quickly degrade evidence of their presence such as tracks or scat. Wildlife biologists attempt to confirm sightings of large carnivores received from the public, and to do so they require reviewable evidence . A visual sighting alone is not enough. Confirmed sightings must be verified by the presence of the animal (in the case of a roadkill for example) or unambiguous game camera images, videos or photos, followed up by a site visit by wildlife biologists to verify location.

When documenting tracks or other signs, it is important to photograph individual tracks as well as groups of tracks. Including a ruler, measuring tape, or an object of standard size (e.g., quarter, business card) in the pictures aid in determination of track size. Including images of the wider area where the tracks were found as well as other local features that can be located if the tracks are compromised by weather is also useful.

Mountain lions are elusive predators, so most sightings last only a few seconds. In Illinois, domestic dogs, domestic cats, and bobcats ( Lynx rufus ) are animals commonly misidentified as mountain lions. Large dog tracks are often misidentified as mountain lion tracks, but there are several key differences. Cats, including mountain lions, walk with their claws retracted, so claw marks will not appear in their tracks. The other differences are highlighted in this graphic of a mountain lion track and a dog track.

An illustration of the difference between a mountain lion footprint and a large dog footprint. The two front toes of a mountain lion are at a slight angle to one another unlike a dog's two front toes which are in line with one another. Mountain lion tracks are also larger with an indented paw pad which has three lobes. A dog track is likely to show claw marks but a mountain lion track will not have claw marks visible.

Mountain lions have been protected in Illinois since 2015, when SB3049 amended the Illinois Wildlife Code by adding mountain lions , black bears and gray wolves to the list of protected species. Because they are protected, mountain lions may not legally be hunted, killed, or harassed unless there is an imminent threat to people or property. Should you see a mountain lion on your property, and you feel that you or your property is being threatened, contact the IDNR to learn about the options available to address potential threats. The IDNR may assist you with control measures.

We should expect dispersing male mountain lions to continue to traverse through Illinois in the future, though biologists expect it to be well into the future before a breeding female establishes a permanent home range in Illinois. This is because female mountain lions rarely disperse long distances in search of new quality habitat or mates.

To learn more about mountain lions, including their basic natural history as well as updated information on confirmed mountain lion sightings, visit the mountain lion page at Wildlife Illinois .

Laura Kammin is a Natural Resources Specialist with the National Great Rivers Research and Education Center. She formerly held positions at Illinois-Indiana Sea Grant, University of Illinois Extension, Prairie Rivers Network, and the Illinois Natural History Survey. She received her master’s degree in wildlife ecology from the University of Illinois, Urbana-Champaign.

Scott F. Beckerman received B.S. and M.S. degrees in Fisheries and Wildlife Management from the University of Missouri-Columbia. He has been helping people resolve conflicts with wildlife since the late 1980s and has been working with the USDA APHIS Wildlife Services program as a wildlife biologist in Missouri, Iowa, Wisconsin and California since 1992. Scott has been the State Director of the Wildlife Services program in Illinois since 2006.

Dakota Bird received a B.S. degree in Wildlife and Fisheries Management from the University of Tennessee. He has held various Large Carnivore and Wildlife Damage Management positions which include Wolf Technician in Minnesota (USGS), Domestic Sheep Protection/Urban Wildlife Specialist in Arizona (USDA WS), and Black Bear Management in Tennessee (NPS) and Wisconsin (USDA WS). Currently, he is a Wildlife Biologist for USDA APHIS Wildlife Services Illinois Program (WS IL) where his primary duty is to serve as the liaison between WS IL and the Illinois Department of Natural Resources to ensure collaboration between the two agencies when it pertains to Large Carnivores and Chronic Wasting Disease.

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Question: I may have seen him on thrusday night about 12:30 am in Hillsboro illinois I can’t be sure i ran in the house but I think so

Question: I believe I saw a mountain lion do you guys want to know about it

Question: Is this a cougar. https://outdoor.wildlifeillinois.org/articles/a-tale-of-two-big-cats-what-we-learned-about-the-mountain-lions-in-illinois-in-2022

Turpentine Creek Wildlife Refuge

Mountain Lion

Did you know? Cougars can be found in both North and South America

Marissa Mountain Lion

Kingdom : Animalia

Phylum : Chordata 

Class : Mammalia 

Order : Carnivora 

Family : Felidae 

Subfamily : Felinae 

Genus : Puma  

Species : concolor  

Scientific Name : Puma concolor  

IUCN Red List Status: Least Concern , but population is decreasing

Habitat : Mountain lions are the widest distributed mammal, ranging from North America to South America. Due to their broad distribution, they are found in a wide variety of habitats such as forests, deserts, and mountains. They tend to prefer rough, rocky terrain with moderately dense, low vegetation. For mountain lions to survive successfully in an area, they have three main requirements: freedom from excessive human interference, dense forest cover for hunting, and adequate prey populations, like large ungulates.

Cougar Species Range Map

Territoriality : Mountain lions are extremely territorial, males will occupy a territory of about 150 square miles. A male will actively protect his territory by continually walking around the perimeter. If another male comes near, the original male will fight to defend his territory. Females will typically occupy about 50 square miles; they will not actively defend and fight, but they do tend to avoid each other.  One male’s boundaries will usually overlap two to three females’ territories, all for the potential for him to breed with them. As young mountain lions mature, females will generally stay close to their mother, but males will roam further away. To avoid inbreeding, a male may travel up to 100 miles away. 

In the Wild: Up to 10 years

In Captivity: Up to 20 years

Estimated 20,000 – 40,000 in the United States

Physical Description

Weight: 80-200 pounds

Length: 7-8 feet (nose to tail)

Mountain lions are the largest member of the cat family in North America. They vary in size and weight, with males being larger than females. Compared to other wild cat species in North America, mountain lions are much larger than lynx or bobcats and have a long tail that measures one-third of their total body size. Mountain lions have short, coarse fur that can be in two color phases- red and gray. The red phase varies from buff, cinnamon, and tawny to a very reddish color. The gray phase varies from silvery gray to bluish and slate gray. Some parts of their body, such as their muzzle and tip of the tail, are black. Newborn mountain lions are heavily spotted for the first three months of their life, but the spots begin to fade soon after. These spots have usually disappeared by six months of age, but it can vary depending on the cat.   

Marissa Mountain Lion

Mountain lions were once found over most of the Western Hemisphere from Canada to the tip of South America, and from the Pacific to the Atlantic oceans. During the time of European settlement into the United States, mountain lion populations saw an enormous decline.  The early settlers feared the cats and believed they had a negative effect on game populations. By the 1800s, these cats were extirpated or severely reduced in the eastern U.S due to the human population increase. Because of this increase, mountain lion habitats declined, and their primary source of food, whitetail deer, saw a significant loss. In the western U.S, mountain lion populations were lowered by the 1900s. By the late 1960s, mountain lions were given game-mammal status in many states, leading to a broader range of population management and programs. After this was enacted, their populations began steadily increasing in numbers. 

Today’s mountain lion population only exists in a fraction of their historic range. While they may not be as populous as they once were, mountain lions can survive in an incredibly diverse array of ecosystems. They currently span 28 countries in North, Central, and South America. Fifteen western states and Florida actively acknowledge breeding populations, while many other states have occasional sightings of the species. 

Reproduction

Gestation: 90 days

Litter size: 1 to 4 (2 to 3 average)

Mountain lions are capable of breeding at any time during the year, but most give birth between April and July, following a three month gestation period. During the breeding season, the infamous scream of the female mountain lion may be heard; this scream is used to attract males. When it is time to give birth, females will find a den in a secure location. An average of 2-3 young, called kittens, are born and rely solely on the mother. She will defend them against males who may try to kill them. Kittens will nurse exclusively for two months; then they will accompany their mother on hunting trips to learn.  Young mountain lions stay with the mom for two years. After this time, young males will travel hundreds of miles away to find a new territory. 

Diet and Hunting Behaviors

A mountain lion is an obligate carnivore, meaning they solely eat meat. Deer are their primary prey, but they are not limited only to deer. They will prey on other ungulates such as elk, moose, and sheep, as well as smaller mammals such as rabbits, porcupines, and skunks. In places where human development has taken over mountain lion habitat, the cats have been known to prey on livestock, domestic horses, dogs, and cats. 

Mountain lions are known as ambush hunters who will wait patiently in dense vegetation or rock crevices. Once their prey has walked by, the cat will silently stalk behind, which results in a quick, surprise attack. They will kill with a powerful bite below the base of the skull, usually breaking the neck. Once their prey is dead, mountain lions will drag the carcass to a sheltered spot, eating 20 to 30 pounds in one sitting. Once they are full, they will bury it under dirt, leaves, or snow to return to later.    

Mountain lions used to roam nearly all of the United States. After becoming a highly sought after hunting trophy and a history of attacking livestock, they were persecuted by both hunters and farmers. By the 1900s, mountain lions were eliminated from nearly all of their range in the Midwest and Eastern United States. Because they exist across the country, some in small, isolated populations, all mountain lions face threats such as trophy hunting, poisoning, predator control, and habitat loss. 

Today, mountain lions are found primarily in the western part of the United States, but have been seen in more eastern states such as Missouri and Arkansas. In 2011, a subspecies of mountain lion, the eastern cougar, was declared officially extinct by the U.S. Fish & Wildlife Service. Florida panthers, another subspecies in the United States, is listed as critically endangered as there are only an estimated 120-230 combined juveniles and adults remaining. Efforts to preserve the species were officially created in the 1960s, but people are not always eager to help predator populations recover. 

Due to the mountain lion’s ability to travel long distances, the cats occasionally wander into areas that may seem inappropriate, including places densely settled by humans. Even though their appearances are almost always extremely brief, public panic can lead to the pursuit of the animal. 

For people who live in or around wildlife habitats, mountain lion sightings and attacks on pets or livestock may occur. Attacks by mountain lions are rare, but there are ways to prevent them from occurring. If you have any questions or concerns about wildlife in your area, contact your local wildlife management officers. 

The conservation of mountain lions depends on the preservation of large amounts of habitat. They require about 13 times as much area as a black bear or 40 times as much as a bobcat to thrive. By preserving enough wilderness to support a stable mountain lion population, countless other species of plants and animals can benefit. 

Mountain Lion Safety

  • Keep your distance, and take this rare encounter very seriously. DO NOT approach the cat 
  • Seem as large as possible, pick up small children, leash pets, and stand close to those around you. Open your jacket and raise your arms, and wave them slowly.
  • Make loud noises that do not sound like a prey animal, such as banging your water bottle or walking stick. Speak slowly and loudly to disrupt the mountain lion’s hunting instincts. 
  • Do not act afraid, act defiant by maintaining eye contact, and DO NOT run away. DO NOT bend over or crouch down. Throw stones or branches and aggressively wave your arms. 
  • Slowly create distance and assess which direction to take. Without turning your back to the mountain lion, do not move towards kittens or prey. Give the lion the time and space to get away from you. 
  • Protect yourself if the mountain lion is to attack you. Protect your neck and throat by using rocks, jackets, garden tools, tree branches, but do not give up.

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Mountain Lion Population in Each State (Current Estimates)

 Reviewed by Samantha Smith B.S. on 02-28-2024

mountain lion travel range

There are 6 species of wild cats in North America, and 2 of those are what we consider “big cats”. The smaller wild cats are the bobcat, lynx, ocelot, and jaguarundis. The big cats of North America are mountain lions and jaguars. In this article we’ll be focusing on the former of those big cats in the United States, the mountain lion. We’re going to look at mountain lion population by state, though many states don’t have a population as they are mainly found in Western U.S. states.

Mountain lions have been more prevalent in the west in the last 100 years and many of the pockets of breeding populations east of the Mississippi River have been wiped out over the decades due to habitat loss and hunting. Having said that, through conservation efforts these big cats are making a comeback in many areas with their ranges and populations growing.

mountain lion travel range

So let’s dive right into the article and have a look at all of the U.S. states that mountain lions call home, as well as those that they don’t.

Mountain lions, cougars, pumas and panthers- what’s the difference?

In North America, you can find mountain lions, cougars, pumas and panthers and the only difference between them is the name that is used for them! In other words, these are all different common names used to refer to the same animal. The scientific name for this species is Puma concolor,  and the many different common names for this species demonstrate the importance of the use of binomial nomenclature . 

The reason why mountain lions have several different common names is because they are (or at least once were) widely distributed across the North America, so these names are due to regional or cultural differences. In this article, we will use these names interchangeably. 

U.S. states with Mountain lion populations

Below we will talk about the mountain lion populations in each U.S. state. We will look at the currently accepted population estimates, and any other notable information and history about cougars in that state.

The majority of U.S. states do not have any population of breeding cougars. Note that many of these states have had sightings over the years, and continue to have them, but that doesn’t necessarily constitute a population in the state.

Mountain lions have been extirpated (considered extinct) from east of the Mississippi river for almost 100 years, however roaming cougars from western states often travel east searching for territory.

Mountain Lion population by state

The following population estimates were taken from state government websites, and other authoritative sources. This list is for breeding mountain lion populations, but some sightings are noted. They are accurate to the best of our knowledge.

mountain lion travel range

Mountain Lions have a status of extirpated in Alabama, there have been no confirmed sightings in the state in over 50 years. There are occasional reports of people who think they saw one, maybe they even took a blurry picture, but those are most likely bobcats. Read more about the status of Mountain Lions in Alabama here .

Alaska is one of the few states where mountain lions are not technically native to and there is currently no breeding population of Mountain Lions in Alaska. However, with populations expanding across western states, there is a chance they could begin to travel into Alaska. There have been a few reports of sightings, with most sightings in southern Alaska, with a few rare sightings in the interior. Often, people who thought they spotted a mountain lion saw another member of the wild cat family, the lynx. Learn more about Mountain Lions in Alaska here .

Mountain lions can be found throughout Arizona, and data suggests the populations are not only stable, but growing. The current estimate is 2,500 – 3,000 throughout Arizona. The most common areas to see mountain lions in Arizona are in places of rocky and mountainous terrain. Hunting is legal, but closely regulated by the state. Find out more about Mountain Lions in Arizona here .

4. Arkansas

There is not currently a known breeding population of mountain lions in Arkansas and there hasn’t been in decades. There have been between 20-25 sightings of mountain lions in the state since 2010, but no evidence to support that these animals are breeding here and not just passing through. However, very recently there was a mountain lion found dead in the state, making it the first verifiable sighting in about a decade . Arkansas has a huge feral hog population, and this abundant prey item might be one of the reasons mountain lions have began to wander there.

5. California

With more than half of the state having prime habitat (foothills and mountains), California has a large population of Mountain Lions. They are currently a protected species in the state and cannot be hunted as game. Until fairly recently, it was believed that the population was about 6,000 individuals in the state, but new data suggests that the more realistic number is closer to 4,500. More information about Mountain Lions in California can be found here .

6. Colorado

Mountain lions have been an important part of the Colorado ecosystem for a long time. The states beautiful wilderness and mountainous terrain are prime mountain lion habitats. There are an estimated 3,800 – 4,400 mountain lions in Colorado. Colorado and California have the highest estimated populations of mountain lions in the United States. 

To learn more about “lion country” in Colorado check here .

7. Connecticut

Despite many people claiming they have seen a mountain lion and calling sightings in to the state authorities, only one has been confirmed. Connecticut’s official position is that mountain lions have been extinct in the state since the late 1800s. The one confirmed case, sometimes referred to as the “ Connecticut Mountain Lion ” happened in June 2011 when an SUV struck and killed a cougar in Milford. A trail of DNA evidence collected over the years shows this particular mountain lion had traveled over 1,500 miles through New York, Michigan, Wisconsin and Minnesota. So while this amazing traveler ended up in Connecticut, it did not originate there.

8. Delaware

While there are some sightings reported each year, there have been no confirmed mountain lions found in Delaware. The population is considered zero with perhaps the occasional transient visitor.

A subspecies of mountain lion can be found in Florida, the Florida panther. They are considered to be the only breeding species of cougar in the eastern part of the U.S. Florida panthers were listed as endangered in 1973. A long term plan was enacted in 2008 to try and protect and grow the population and there have been many grassroots efforts to try to protect what’s left of their range. At present they are only found around the Caloosahatchee River / southwest of lake Okeechobee in southern Florida. Their population is thought to be fewer than 100 . Florida Fish & Wildlife has a page with several resources about the Florida Panthers here .

10. Georgia

According to the state wildlife division, there have only been 3 credible mountain lion sightings in the last 25 years, all of which were thought to be related to the Florida panther.  People report sightings in Georgia every year, however the state is not considered to have a population and it is believed most of these sightings are a case of “mistaken identity”. Find more information about Georgia’s stance on mountain lion sightings here .

Being the isolated chain of islands that it is, Hawaii is not home to any mountain lions, or any larger mammals for that matter. There were a handful of questionable sightings in 2002 that drummed up attention, but no big cat was ever found. Some locals call this the “phantom cat” and liken it to the Loch Ness monster. If there were a big cat on the loose, it would have had to escape captivity.

Mountain lions are found regularly throughout Idaho, and have been observed in nearly every county. Estimates place the mountain lion population in Idaho to be around 2,ooo .  It is legal to hunt mountain lion in Idaho. In fact, in a controversial move, the state removed hunting quotas for mountain lions state wide. This decision raises important questions about what the state’s plans are moving forward to monitor their populations to ensure that they are not overhunted. Read more about mountain lions in Idaho here.

13. Illinois

It is not believed that there is currently any breeding population of mountain lions in the state of Illinois. However there are confirmed sightings, so it is likely the big cats do occasionally move through the state. Most recently, a mountain lion was hit and killed on Interstate 88 in Delkalb county that was likely from South Dakota. It is possible numbers in Illinois could slowly increase if cougars in areas such as South Dakota and the Rocky Mountain states begin to disperse eastward. Find out more about mountain lions  in Illinois here .

14. Indiana

Indiana does not have a mountain lion population, although there are occasional sightings. The last sighting reports considered to be confirmed were in 2009-2010. To see Indiana’s Department of Natural Resources pages on mountain lions, check here .

Like many other states we have mentioned, Iowa does not have a breeding population of mountain lions, but still sightings are made every year. A handful of these have been confirmed over the years, but according to the Iowa Department of Natural Resources, 95% of reported sightings are mistaken identity. The IDNR does admit that because of the confirmed cases, it is possible there are a few “free ranging mountain lions” present in the state traveling from nearby states. Read Iowa’s mountain lion pamphlet here .

Kansas is another central U.S. state where there is no established breeding population, but a fair number or mountain lions have been seen over the years. According to the Kansas Department of Wildlife, Parks and Tourism it is likely the sightings are transient young males from more established populations (such as neighboring Colorado). Young males may travel long distances looking for new territory. It is thought these mountain lions only pass through Kansas. For more on Kansas mountain lion sightings, check this page .

17. Kentucky

Kentucky is not home to mountain lions. The nearest wild populations are also many hundreds of miles away in Nebraska and Florida. While reports of sightings have gone up, the Kentucky Department of Fish & Wildlife believes this is due to people actually seeing bobcats, which are increasing in population throughout Kentucky. For more information see the Kentucky DFW page on mountain lions here .

18. Louisiana

There is no breeding population of mountain lions in Louisiana. They are considered to be rare in the state, with only a few confirmed sightings. For example, a cougar confirmed on a trail cam in 2016 was the first confirmed mountain lion in the state in over five years. Over the years there have been some local bigfoot-like folklore about Louisiana black panthers, however the Louisiana Wildlife and Fisheries Department have of course denied these claims.

While mountain lions could once be regularly found in Maine, there has been no population since the early 1900s. There are occasional sightings but any confirmation is rare.

20. Maryland

Bobcats are the only official populous and breeding “big cats” found in the state of Maryland . Like most eastern states, Maryland was once home to the eastern mountain lion but none remain today. There are occasional sightings, and there may possibly be the rare traveler that passes through. The Susquehannock Wildlife Society wrote an article about a potential cougar photograph submitted to them and discussed their process of investigation, in the end they deemed it was not a mountain lion.

21. Massachusetts

Like all New England states, Massachusetts no longer has a population of mountain lions. Only two potential incidents have been confirmed, and both involved finding evidence of mountain lions, not the mountain lions themselves. The most recent being a photo of tracks in snow taken in 2011, and the state believes these may have been made by the “Connecticut Mountain Lion” confirmed in the area during that same time period. Read more about Massachusetts findings on mountain lions here .

22. Michigan

Michigan no longer has an established mountain lion population, however confirmed sightings are starting to become more frequent. Since 2008, there have been 82 confirmed sightings /instances of mountain lions in Michigan. This is still a very small number, but does suggest that mountain lions are traveling slightly more frequently to Michigan from established mountain lion territory in areas like South Dakota, Wyoming and Nebraska. To see the the Michigan Department of Natural Resources page on cougars, visit here .

23. Minnesota

Minnesota does not have an established breeding population of mountain lions. However, it boarders North and South Dakota, both of which do have established mountain lion populations. Therefore roaming cats moving west are expected to sometimes cross into Minnesota. The state department of natural resources says that there is evidence that the prevalence of mountain lions in the state is increasing, however they believe it is still mostly just animals wandering in and out, not staying to breed. For more on cougars in Minnesota, see the MDNR page here .

24. Mississippi

After extirpation over 100 years ago, mountain lions have not returned to Mississippi. There is currently no breeding population and reports are incredibly rare and most often unsubstantiated.

25. Missouri

Missouri does not currently have a breeding population of mountain lions, however the Missouri Department of Conservation acknowledges that mountain lions do have a presence in the state. They even have a “ Mountain Lion Response Team ” to investigate sightings and gather evidence to try and track the population. Almost all of the mountain lions found in Missouri so far, at least those where a sex determination could be made, have been males. Males are known to sometimes wander far in search of their own territory. Between 1994 and late 2023, there have been 117 confirmed reports. One of the more recent, credible reports was made when an elk was found that was undoubtedly killed by a mountain lion. For more on Missouri’s mountain lion reports check here .

26. Montana

After an unfortunate past of over hunting, Montana’s mountain lion population has made a comeback.  Starting in 1879 and lasting many decades, bounties were paid for killing mountain lions in Montana, severely impacting the population. In 1971 they were classified as “game animals”, and hunting could therefore be controlled by the state. This plus increasing numbers of deer and elk allowed the lion population to slowly recover.

The Montana Fish, Wildlife and Parks department conducts in depth population monitoring of two of it’s four designated ecoregions. In these two ecoregions, the estimated population is at least 2,500 , but the total population for the entire state is unknown. Learn more about mountain lions in Montana here .

27. Nebraska

Mountain lions were eliminated from Nebraska by the late 1800’s, and it wasn’t until the 1990’s when they began to return. There are currently three areas of Nebraska where the mountain lions are living and breeding, Pine Ridge, Niobrara River Valley and Wildcat Hills. As of 2021 the population in Pine Ridge was estimated at 33. The Niobrara River Valley and Wildcat Hills populations are relatively new with an estimated minimum of 20 individuals in the Niobrara River Valley and no estimate for the Wildcat Hills yet. The Nebraska Games & Parks Commission is committed to tracking and controlling the population to keep it an an optimal level. To read more about their efforts and mountain lions in Nebraska visit their page here .

It is thought that mountain lions were once relatively rare in Nevada, but as the populations of deer grew in the 1930’s and 40’s, so did the cougars. Estimates seem to fluctuate but the most recent I could find approximates about 3,400 mountain lions . According to the Nevada Department of Wildlife “In Nevada lions are found in areas of pinion pine, juniper, mountain mahogany, ponderosa pine and mountain brush. Lions generally are most abundant in areas where deer are plentiful.” Nevada has a brochure about living with mountain lions here .

29. New Hampshire

The mountain lion population in New Hampshire is considered to be zero. According to the New Hampshire Fish and Game page, there continues to be no evidence of any mountain lions in the state.

30. New Jersey

Officially, the bobcat is the only large cat that calls New Jersey its home, no mountain lions. However like most eastern states, there are occasional sightings called in, however none that we could find that have been confirmed.

31. New Mexico

The most recent estimate from a 2023 report puts the population of mountain lions in New Mexico at 3,494 . The state has begun to use more high tech methods for tracking these secretive lions, including motion-capture cameras, mathematical models and GPS collaring. This research will contribute to a better understanding of mountain lion ecology and can help inform conservation efforts. They also engage in harvesting specific numbers of cougars each year to control the population. Read more at the New Mexico Game & Fish cougar page here .

32. New York

There is no population of mountain lions in New York state. But there have been sightings. According to the states’ environmental conservation page, none of these were native and breeding in NY. One was confirmed to have traveled about 1,800 miles from a South Dakota population, and a few others were cougars that escaped from a licensed NY facility. You can read more about how to report a mountain lion sighting in New York here .

33. North Carolina

There are no known mountain lions in North Carolina and no credible sightings have been made since 1886. Like many other states, there are sightings reported each year to the state, but no confirmations have been made.

34. North Dakota

North Dakota is thought to be one of the most recent states to be colonized by mountain lions as their populations have begun to expand. While much of North Dakota is open prairie habitat that will not support many mountain lions, they do reside in the more mountainous western parts of the state.  The state estimates that as of 2023, there are approximately 50 individuals in North Dakota. Despite their small numbers, the hunting quotas for mountain lions are relatively large which draws questions about how sustainable these quotas are. 

While there are many unconfirmed sightings of mountain lions in Ohio, there is not believed to be any living in the state. In fact an article I read noted that a confirmed mountain lion sighting hadn’t happened in over 100 years. The cat that is growing in population within Ohio is the bobcat, and many official believe most of the sightings called in are actually people mistakenly seeing bobcats.

36. Oklahoma

Mountain lions are considered to be a “transient species” in the state of Oklahoma. Meaning they have been documented in small numbers in the state, but there is no evidence that they breed or have established a territory in any one area. Visit Oklahoma’s mountain lion research page here .

Oregon has quite a healthy population of mountain lions, estimated to be over 6,000 however only about half of these individuals are thought to be breeding adults. They can be found through the state, with the highest populations in the Blue Mountains and Cascade Mountains. Despite these large numbers, they stay in the wilderness and encounters with humans have been rare. Visit Oregon’s mountain lion page here .

38. Pennsylvania

Pennsylvania currently does not have a population of mountain lions. Occasional sightings are made, but nothing confirmed. There is a possibility a transient lion passes through the state from time to time, but this would be considered rare.

39. Rhode Island

There is no known population of mountain lions in Rhode Island. Like most New England states there are always sightings, but currently no confirmations.

40. South Carolina

According to the South Carolina Department of Natural Resources, the state does not have a free-ranging wild cougar population. They do acknowledge some have escaped captivity in the last 30 or so years, and that there is a possibility of the occasional transient young male passing through. Some believe the Florida panther has traveled up to South Carolina but there is currently no evidence of that.

41. South Dakota

Most of the mountain lions in South Dakota can be found in the Black Hills region, where there are an estimated 200-300 lions, according to a 2023 report . The report states that the population is stable for now, but if the mortality rates stay the same, then it will be difficult for the population to remain stable. There are also mountain lions in other parts of the state but the populations are smaller and exact numbers unclear.

42. Tennessee

Only a few confirmed sightings of mountain lions have occurred in Tennessee in recent years. They are considered a transient species in the state, with no breeding population. You can find out more about the confirmed sightings on the Tennessee Wildlife Resources Agency page here .

mountain lion travel range

Map from tpwd.texas.govTexas Parks and Wildlife put together a nice PDF about the state’s mountain lions here .

Utah is home to approximately 2,300 mountain lions, which is a slight decrease from previous years. This slight decline is thought to be a result of several serious droughts which has impacted populations of a major prey source for mountain lions, mule deer . Currently, hunting is allowed, with the state controlling the number of allowed harvests every year. You can read the states 2015-2025 cougar management plan here .

45. Vermont

Mountain lions are considered to be extirpated from the state of Vermont. They once populated the state, colloquially called the “Catamount”. The catamount remains the mascot for the University of Vermont to this day. The last catamount / mountain lion in Vermont was killed in 1881. Occasional sightings persist, but rarely is there confirmation.

46. Virginia

There is no breeding population of mountain lions in Virginia, with the last wild cougar in the state killed in Washington County in 1882. Experts say there is habitat for them in the Appalachians today, and as the western mountain lions continue to slowly roam to the east, they could one day return more permanently to Virginia.

47. Washington

According to a 2022 report from the Washington Department Fish and Wildlife, the mountain lion population is approximately 3,600. They are distributed throughout the state, except for a pocket around the Columbia river basin where not many reside. As the population of humans in these areas, especially western Washington, increases, encounters may become more common. However cougar attacks remain rare, with only two fatal attacks in the past 100 years. Visit the Washington cougar page here .

48. West Virginia

Just like their neighbor Virginia, there is no breeding population of mountain lions in West Virginia. There continue to be sightings and it is likely a transient mountain lion roams through on rare occasions.

49. Wisconsin

There is no permanent breeding population of mountain lions in Wisconsin, however there are occasional confirmed sightings. Most recently, a bowhunter shot and killed a male mountain lion when he feared that the animal was going to attack him. Biologists believe that the mountain lions seen belong to the groups found out in the Black Hill of South Dakota.

50. Wyoming

It is estimated there are about 2,000 mountain lions in Wyoming. In recent years the population of certain areas has decreased dramatically, due to a “perfect storm” of reintroduction of gray wolves, increased hunting of elk, and increased hunting of the mountain lions. There is a brewing debate in the state of whether mountain lion hunting should be taken off the table until population numbers rebound.

About Mountain Lions

mountain lion travel range

Mountain lions were historically found throughout nearly all of North and South America. Indigenous peoples of both continents often revered and had much admiration and folklore surrounding the big cats. As European settlers arrived and populations grew, cougars became a prized animal to hunters, and a hated threat to farmers and ranchers livestock.

Through hunting and habitat loss, mountain lions were completely wiped out of the eastern United States and faced harsh population declines in others by the early 1900s. Due to changes in laws, protections and conservation efforts, mountain lion populations have rebounded in some western states. Many experts believe with the high deer population (cougars favorite prey animal) that mountain lions could begin to make a comeback in their historic eastern range, given enough available habitat.

Mountain lions are very adaptable to different types of environments, which is why they are so widespread across North and South America. They can live in many types of forest, as well as canyonland, lowland and mountainous deserts, dense brush and even open spaces with little cover. They don’t form packs though, preferring to stay solitary, and only a few can exist together peaceably within a 30 square mile range. This makes how much suitable land is available for them to roam one of the biggest limiting factors in their population.

Because cougars are so adaptable to different types of habitat, they have a huge range. From the northern Yukon in Canada, down to the southern Andes at the bottom of South America. In the United States, they was extirpated from most of the states east of the Mississippi River in the 200 years after European colonization. The U.S. range today is mainly across the western states with a small population in southern Florida. There is evidence that these western mountain lions are slowly spreading east and may recolonize the middle and eastern states.

mountain lion travel range

The favorite meal of mountain lions are deer and elk. They hunt by stealth, often at dawn/dusk or at night, and often will surprise their prey from behind in an ambush.  An adult cougar will consume an average of one deer per week. They will also hunt feral hogs, raccoons , rodents, porcupines and even coyotes. When they take prey that is too large to eat in one sitting, they will often drag it off to a protected spot and cover it in pine needles, grass or dry leaves to keep it hidden from other predators or scavengers that might want to steal their meal. This behavior may also slow down spoilage of the meat. They can then come back several times during the week to feed on the carcass.

Mountain lions are the fourth largest cat species in the world, behind the lion, tiger and jaguar.

  • Length (nose to tail tip): Males 7.9 ft, Females 6.7 ft. Tails account for about 25-35% of their body length.
  • Height (at the shoulder): 24-35 inches
  • Weight: Males 117-220 pounds, Females 64-141 pounds

Mountain lions at the poles are larger than those found at the equator. Coat color can vary with location also, sandy brown, reddish brown, to silvery – light gray. Whatever their color, it remains uniform across their body with no spots, stripes or patterns.

Many animals have a “breeding season” and only breed during certain parts of the year. However mountain lions can breed year round. Males and females don’t spend time together outside of the few day period when they decide to breed. The typically silent cats will make loud screaming noises. It is thought these sounds are made by the females to indicate they are in heat and ready to mate, while if the male screams it is to get the attention of females over other male rivals.

Females usually only give birth once every two years, and the litter size is typically three cubs, but can range from one to six. Cubs are born after 90 days, born small and blind, kept hidden by mom in thick vegetation for about 40-70 days. Males are not involved in the lives of the cubs after they are born. The cubs will continue to stay with their mother until about 10-26 months old. Sadly, it is common for at least one cub in the litter to die before reaching two years of age. But for those that make it to 48 months, they begin to disperse in search of their own home range.

While all mountain lions fall under the species Puma concolor, there has long been debate on how many subspecies exist across the cougars vast range. At one point there were over 30, but those began to narrow as DNA testing became possible and many were found to not be genetically different enough to be considered separate subspecies. In 2005, six subspecies were recognized:

  • P.c. concolor
  • P.c. cougar
  • P.c. costaricensis
  • P.c.anthonyi
  • P.c.cabrerae

Depending on who you talk to, some scientists consider that only a few of those are valid distinctions. This list will likely continue to evolve as more sophisticated and widespread DNA testing is performed.

Another helpful resource for state cougar information is the cougarfund.org

mountain lion travel range

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Cougar (Puma concolor)

 Adult male cougar stands in tree.

Sleek and graceful, cougars ( Puma concolor ) are solitary and secretive animals rarely seen in the wild. Also known as mountain lions or pumas, cougars are known for their strength, agility, and awesome ability to jump.

WDFW estimates there are approximately 3,600 cougars in Washington state.

Description and Range

Physical description.

Cougars are the largest members of the cat family in Washington. Adult males average approximately 140 pounds but in rare cases may weigh 180 pounds and measure 7-8 feet long from nose to tip of tail. Adult males stand about 30 inches tall at the shoulder. Adult female cougars rarely exceed 110 pounds. Cougars vary in color from reddish-brown to tawny (deerlike) to gray, with a black tip on their long tail. Cougar kittens are spotted until they are 4-5 months old, after that, barring patterns may remain up to 14 months of age. For additional information, see this cougar brochure .

Geographic range

Cougars occur throughout Washington where suitable cover and prey are found. Cougars use steep canyons, rock outcroppings and boulders, or vegetation -- such as dense brush and forests -- to remain hidden while hunting. Adult male cougars roam widely, covering a home range of 50 to 150 square miles, depending on the age of the cougar, the time of year, type of terrain, and availability of prey. Adult male cougars’ home ranges will often overlap those of three or four females. Female home ranges are about half that of males and there in considerable overlap in female home ranges. Often female progeny will establish a territory adjacent to mother, while virtually all males disperse considerable distances from the natal area. Cougar's range often overlaps with other predators like wolves and bobcats. WDFW is currently conducting the Washington Predator-Prey Project that looks at the impact of wolf recovery on other species like cougars, deer, and elk. 

Regulations

Licenses and permits.

A valid big game license, which includes cougar as a species option, is required.

Rules and seasons

General seasons.

Early hunting season: Sept. 1 - Dec. 31

Late hunting season: Jan. 1 - Apr. 30 or when the harvest guideline is reached, whichever occurs first

Harvest guideline system

In hunt areas with a harvest guideline, the cougar late hunting season may close on or after January 1 in one or more GMUs if cougar harvest meets or exceeds the guideline.

Starting Jan. 1, hunters may hunt cougar until the hunt area harvest guideline has been reached, or April 30, whichever occurs first.

All cougars killed by licensed hunters during the early and late hunting seasons, and seasons authorized by the Fish and Wildlife Commission shall be counted toward the harvest guideline.

It is each cougar hunter’s responsibility to verify if the cougar late hunting season is open or closed in hunt areas with a harvest guideline. Cougar hunters can verify if the season is open or closed by calling the toll free cougar hunting hotline (866-364-4868) or visiting the department's website . The hotline and website will be updated weekly beginning Jan. 1.

One cougar per license year.

It is unlawful to kill or possess spotted cougar kittens (usually <80 lbs ) or adult cougars accompanied by spotted kittens. Since cougars are solitary, observing multiple tracks suggests a female with offspring. However, smaller kittens may not be visible to hunters so please observe and be patient before shooting.

Hunting methods

Hunters may use any legal weapon for hunting cougar.

Hunting with dogs

The use of dogs for recreational harvest of cougar is prohibited statewide.

Mandatory reporting and hide-sealing requirements

All successful cougar hunters must report cougar harvest to the cougar hotlines at 866-364-4868 within 72 hours (state holidays are not counted as part of the 72 hours) and must have the pelt sealed within five days of notification. Contact a WDFW office to set up an appointment to have the pelt sealed. Ensure proof of sex is left attached to the pelt.

The skull and hide (both non-frozen) must be presented so teeth and biological samples can be taken. Placing a stick or wedge in the cougar's mouth after harvesting it aids in the removal of teeth.

No one may possess an open WDFW cougar seal unless it has been cut by a licensed taxidermist or fur dealer who has received and invoiced the pelt for processing (WAC 220-415-050).

See the  Cougar hunting area openings and closures page for the current status of cougar hunt areas.

Living with wildlife

Food and feeding habits.

Image of a cougar skull.

Cougars are most active from dusk to dawn, but it is not unusual for cougars to hunt anytime during the day. Adult cougars typically prey on deer, elk, moose, mountain goats, and wild sheep, with deer being the preferred and most common prey. Other prey species, especially for younger cougars, include raccoons, coyotes, rabbits, hares, small rodents, and occasionally pets and livestock (e.g. goats, sheep, and chickens). A large male cougar living in the Cascade Mountains kills a deer or elk every 9 to 12 days, eating up to 20 pounds at a time and burying the rest for later. Except for females with young, cougars are lone hunters that wander between places frequented by their prey, covering as much as 15 miles in a single night. Cougars rely on short bursts of speed to ambush their prey. A cougar may stalk an animal for an hour or more.

Feeding areas (caches)

Cougars usually carry or drag their kills to a secluded area under cover to feed, and drag marks are frequently found at fresh kill sites. After killing a large animal and having eaten its fill, a cougar often will cover the remains with debris such as snow, grass, leaves, sticks, or soil. Even where little debris is available, bits of soil, rock, grass or sticks may be used to cover the carcass. The cougar may remain in the immediate vicinity of its kill, guarding it against scavengers and eating it over a period of six to eight days. (Meat becomes rotten quickly in the summer and male cougars have to patrol their territory. Often these males will make a kill, feed until full, leave to patrol the area, and return to feed on the carcass days later.)

Do not approach or linger around a recently killed or partially covered deer or elk.

Cougar tend to leave “soft” tracks, meaning the animals make very little impact on the ground, and their tracks may be virtually invisible on packed earth or crusted snow. In addition, to preserve their sharpness for gripping prey, these animals keep their claws retracted most of the time, and so claw marks are rarely visible in their tracks.

Because cougars carry their heavy tail in a wide U shape at a normal walk, in snow, the lowermost portion may leave drag marks between each print.

Cougars generally cover their droppings with loose soil. When visible, their droppings typically resemble those of most species in the dog and cat families. However, cougars have well developed premolars that can slice through bone and hide. Therefore, their droppings often show chunks and fragments of chewed bone and considerable hair from the hide. Members of the dog family gnaw on bones but usually don’t chew them up into cut fragments.

Drawing of a cougar

Cougar droppings are generally cylindrical in shape, segmented, and blunt at one or both ends. An average dropping measures 4 to 6 inches long by 1 to 1½ inches in diameter. The size of the dropping may indicate the size of the cougar.

Daybed sites

A cougar’s daybed is used for rest, protection from the weather, and to raise young. Cougars don’t use dens like bears do. They may settle down for up to six weeks while the kittens are immobile, but afterward are almost always on the move, making daybeds as they go. In rough terrain, daybeds are usually in a cave or a shallow nook on a cliff face or rock outcrop. In less mountainous areas, day beds are located in forested areas, thickets, or under large roots or fallen trees. Daybeds are frequently near kill sites. No day beds preparation takes place.

Scratching posts

Like house cats scratching furniture, cougars mark their territory boundaries by leaving claw marks on trees, stumps, and occasionally fence posts. Claw marks left by an adult cougar will be 4 to 8 feet above the ground and consist of long, deep, parallel scratches running almost vertically down the trunk. These gashes rarely take off much bark; tree-clawing that removes much bark is probably the work of a bear. (Bobcat claw marks are normally 2 to 3 feet above the ground; domestic cat scratching occurs at a height of about 1½ to 2 feet).

Cougars hiss, purr, mew, growl, yowl, chirp, and cry. The most sensational sounds they make are the eerie wailings and moans heard at night during mating season, especially when competing males have intentions toward the same receptive female. Such wails have been likened to a child crying, a woman’s scream, and the screeching of someone in terrible pain.

Preventing conflict

The cougar’s ability to travel long distances occasionally brings these cats into seemingly inappropriate areas, even places densely settled by humans. Such appearances are almost always brief, with the animal moving along quickly in its search of a suitable permanent home. However, when humans occupy areas in or near wildlife habitat, and people choose to keep livestock animals such as goats, sheep, and chicken cougar sightings and attacks on livestock and pets may occur.

Cougar attacks on humans are extremely rare. In Washington state, there have been two fatal cougar attacks and approximately 20 other recorded encounters that resulted in human injury in the last 100 years.

If you are living in cougar country, prevent a conflict with them by using the following management strategies around your property, and, if possible, encourage your neighbors to do the same.

Modify the habitat around your home. Light all walkways after dark and avoid landscaping with plants that deer prefer to eat. Where a deer goes, a cougar may follow. Shrubs and trees around kids’ play areas should be pruned up several feet to prevent cougars from hiding behind them.

Although costly and not 100 percent effective, a chain-link or heavy woven wire fence that is 10 feet high with 3-foot extensions installed at a 65-degree angle on each post may keep cougars out of an enclosed area. To increase effectiveness, string barbed wire or four electric wires between the extensions, alternating positive and negative wires.

Don’t feed wildlife and feral cats (domestic cats gone wild). This includes deer, raccoons, and other small mammals. Remember predators follow prey.

Close off open spaces under structures. Areas beneath porches and decks can provide shelter for prey animals.

Feed dogs and cats indoors. If you must feed outside, do so in the morning or midday, and pick up food and water bowls, as well as leftovers and spilled food, well before dark. Pet food and water attract small mammals that, in turn, attract cougars.

Keep dogs and cats indoors, especially from dusk to dawn. Left outside at night, small dogs and cats may become prey for cougars.

Use garbage cans with tight-fitting lids. Garbage attracts small mammals that, in turn, attract cougars.

Livestock and small animals, such as goats, sheep, and chickens, are attractants to cougars. Outdoor livestock should be kept in secure pens and away from forest boundaries. For a larger property, consider using a guard animal. There are specialty breeds of dogs that can defend livestock. As with any guard animal, pros and cons exist. Purchase a guard animal from a reputable breeder who knows the animal he or she sells. Some breeders offer various guarantees on their guard animals, including a replacement if an animal fails to perform as expected.

Download our Discover Washington's cougars brochure .

Download our Coexisting with cougars in Washington: a guide for small livestock owners.

Coexistencia con los pumas en Washington: guía para pequeños granjeros

与 Washington 的美洲狮 和谐共存: 告小型牲畜主人知

Cùng tồn tại với báo sư tử ở Washington Hướng dẫn dành cho các chủ chăn nuôi nhỏ

Encountering a cougar

Relatively few people will ever catch a glimpse of a cougar much less confront one. If you come face to face with a cougar, your actions can either help or hinder a quick retreat by the animal.

Here are some things to remember:

  • Stop, pick up small children immediately, and don’t run. Running and rapid movements may trigger an attack. Remember, at close range, a cougar’s instinct is to chase.
  • Face the cougar. Talk to it firmly while slowly backing away. Always leave the animal an escape route.
  • Try to appear larger than the cougar. Get above it (e.g., step up onto a rock or stump). If wearing a jacket, hold it open to further increase your apparent size. If you are in a group, stand shoulder-to-shoulder to appear intimidating.
  • Do not take your eyes off the cougar or turn your back. Do not crouch down or try to hide.
  • Never approach the cougar, especially if it is near a kill or with kittens, and never offer it food.
  • If the cougar does not flee, be more assertive. If it shows signs of aggression (crouches with ears back, teeth bared, hissing, tail twitching, and hind feet pumping in preparation to jump), shout, wave your arms and throw anything you have available (water bottle, book, backpack). The idea is to convince the cougar that you are not prey, but a potential danger.
  • If the cougar attacks, fight back. Be aggressive and try to stay on your feet. Cougars have been driven away by people who have fought back using anything within reach, including sticks, rocks, shovels, backpacks, and clothing—even bare hands. If you are aggressive enough, a cougar will flee, realizing it has made a mistake. Pepper spray in the cougar’s face is also effective in the extreme unlikelihood of a close encounter with a cougar.

Professional assistance

Wildlife offices throughout Washington respond to cougar sightings when there is a threat to public safety or property. Cougars involved in human conflict may be live-trapped by trained fish and wildlife personnel and moved to more remote areas, or removed.  However, both are expensive, time consuming, and do not address the root cause of the problem. It is common that husbandry practices need to be addressed to avoid a repeated conflict.

Contact your local wildlife office for additional information, and in the case of an immediate emergency, call 911 or any local law enforcement office, such as the state patrol.

Cougars and kids

  • Talk to children and teach them what to do if they encounter a cougar.
  • Encourage children to play outdoors in groups, and supervise children playing outdoors.
  • Consider getting a dog for your children as an early-warning system. A dog can see, smell, and hear a cougar sooner than we can. Although dogs offer little value as a deterrent to cougars, they may distract a cougar from attacking a human.
  • Consider erecting a fence around play areas.
  • Keep a radio playing when children are outside, as noise usually deters cougars.
  • Make sure children are home before dusk and stay inside until after dawn.
  • If there have been cougar sightings, escort children to the bus stop in the early morning. Clear shrubs away around the bus stop, making an area with a 30-foot radius. Have a light installed as a general safety precaution.

Precautions for hikers and campers

Drawing of two backpackers seen from behind

While recreating in a cougar’s territory, you can avoid close encounters by taking the following precautions:

  • Hike in groups and make enough noise to prevent surprising a cougar.
  • Avoid hiking after dark.
  • Keep small children close to the group, preferably in plain sight ahead of you.
  • Do not approach dead animals, especially recently killed or partially covered deer and elk.
  • Be aware of your surroundings, particularly when hiking in dense cover or when sitting, crouching, or lying down. Look for tracks, scratch posts, and partially covered droppings.
  • Keep a clean camp. Reduce odors that might attract mammals such as raccoons, which in turn could attract cougars. Store meat, other foods, pet food, and garbage in double plastic bags.

Public health concerns

Cougars rarely carry any communicable diseases that are regarded as threats to humans in Washington.

Feline distemper ( Feline panleukopenia ) antibodies have been documented in Washington cougar populations, but the degree that the Feline panleukopenia virus causes cougar mortality, or is transferred to domestic cats, is unknown.

Legal status

Because the legal status, hunting restrictions, and other information relating to cougars change, contact your local wildlife office for updates.

Cougars are classified as game animals and an open season and a hunting license are required to hunt them ( WAC 220-400-020 ). A property owner or the owner’s immediate family, employee, or tenant may kill a cougar on that property if it is damaging domestic animals ( RCW 77.36.030 ). No permit is required.

The killing of a cougar in self-defense, or defense of another, should be reasonable and justified. A person taking such action must have reasonable belief that the cougar poses a threat of serious physical harm, that this harm is imminent, and the action is the only reasonable available means to prevent that harm.

The body of any cougar, whether taken under the direct authority of RCW 77.36.030 , or for the protection of a person, remains the property of the state and must be turned over to the Department of Fish and Wildlife immediately.

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CDFW

  • Conservation

Mountain Lions in California

adult mountain lion profile

Introduction

The California Department of Fish and Wildlife is tasked with ensuring viable mountain lion ( Puma concolor ) populations persist throughout the state where suitable habitat and other environmental conditions allow. Mountain lions are classified as a 'specially protected' species in California following the passage of the The California Wildlife Protection Act of 1990 (Proposition 117).

Statewide mountain lion population estimates are considered stable based on the best scientific knowledge, research, and methods available. CDFW is currently finalizing a statewide population abundance and density estimate for mountain lions using information gained from rigorous statistical analyses using non-invasive scat survey and satellite-collar data. Estimating population densities an elusive species in a state as geographically large and diverse as California is a complex task. Historic studies over the last 40 years have estimated population densities for different habitat types throughout California.

Increased sightings and reports of mountain lions are likely due to the increased presence of home security cameras (e.g., ring doorbells), social media, and personal trail cameras often used for hunting, wildlife photography and leisure purposes.

General Biology

Mountain lions are the second largest felid in North America with the largest geographic range of carnivores in the western hemisphere that stretches from southern Chile to the Canadian Yukon. The mountain lion is known by over 40 different common names including but not limited to puma, cougar, panther, red tiger, catamount, and screamer. Mountain lions inhabit diverse habitat types across California including temperate redwood forest, coniferous / deciduous forest, coastal chaparral, foothills and mountains. They can be found wherever native or introduced ungulates such as mule deer, elk, bighorn sheep, or feral hogs are present.

  • Body Length: 37.5-55 in (female) 42.1-66.1 in (male)
  • Tail Length: 22.4-36.2 in
  • Weight: 50.7-110 lb (female), 85-170 lb (male)
  • Mountain Lion Conservation Program
  • Science and Research
  • Human - Mountain Lion Conflict

Laws and Regulations

  • Recreating in Mountain Lion Habitat

In California, mountain lions are a specially protected non-game species; following the passage of the California Wildlife Protection Act of 1990 (Proposition 117). Mountain lions have not been hunted in California since 1972. The Department strives to conserve mountain lion populations for their ecological and intrinsic values. We have been studying mountain lion populations in the state for more than 40 years and formally established the mountain lion conservation program in 2015 to coordinate statewide mountain lion research and population monitoring, conflict mitigation, and to inform habitat conservation and protection.

The mountain lion conservation program is based in the wildlife health laboratory in the Wildlife Branch of CDFW.

  • Jason Lombardi , Ph.D. - Statewide Large Carnivore Research Coordinator (Lead Mountain Lion Scientist)

California Endangered Species Act Petition

Over the last decade, research has indicated there is a lack of genetic diversity in specific parts of California (i.e., southern California and central coast; Ernest et al. 2014, Gustafson et al. 2019) as a result of human population growth and barriers that restrict connectivity with other mountain lion populations. The Department is pursuing many actions toward managing mountain lions, including coordination with federal, state, and non- governmental organization partners on projects to improve habitat connectivity, and to increase public outreach and education.

In July 2019, the Center for Biological Diversity and the Mountain Lion Foundation petitioned the Fish and Game Commission to list mountain lions as a candidate species under the California Endangered Species Act ( CESA ) within a proposed evolutionarily significant unit (ESU) located in Southern California and the central coast of California. In April 2020, the Commission found that listing of this ESU may be warranted and designated mountain lion within the ESU as a candidate species. As a result, CDFW is now completing a status review of mountain lions within the proposed ESU. At the end of the review, CDFW will make its recommendation on listing to the Commission. Under CESA, species classified as a candidate species are afforded the same protection as listed species. As a result, mountain lions in this proposed ESU are CESA-protected during the review period.

Statewide Scientific Research

The California Department of Fish and Wildlife (CDFW) established a mountain lion conservation program in 2015 to coordinate statewide mountain lion research and population monitoring, and to inform habitat conservation and protection. The department conducts these statewide projects independently and in collaboration with external collaborators from 8 different academic institutions, 3 non-governmental organizations (NGOs), and 5 governmental agencies. Further, these collaborations have been advised by the California Mountain Lion Scientific Working Group, which is a consortium of the leading mountain lion researchers in the state.

Over the last 8 years there has been a focus to address the following questions 1) develop habitat use models of pumas statewide and in all the eco-regions of the state where mountain lions exist; 2) understand puma demographics and develop tools for estimating mountain lion abundance/density statewide and in the ecoregions long-term; 3) quantify genetic variation and gene flow between mountain lion populations in various regions of the state; 4) identify barriers to and travel corridors that may facilitate mountain lion gene flow between various populations; 5) assess prevalence of disease and other factors that might limit health of individuals and populations of mountain lions; 6) understand ecological relationships between mountain lions and co-occurring prey and predators; 7) research trends in and methods for mitigating mountain lion-human conflict, and 8) addressing human dimension issues related to mountain lions.

Statewide research efforts are lead and coordinated by Statewide Large Carnivore Research Coordinator Dr. Jason Lombardi .

Data collected in statewide or regional mountain lion studies are often published in peer-reviewed scientific and scholarly journals. These efforts help guide conservation and management of mountain lions in California and across their geographic range. For more information and list of publications, please check out Mountain Lion Research Publications

External Mountain Lion Scientific Collection Permitted Research

CDFW is responsible for permitting and overseeing mountain lion research in the State.

Qualified individuals, educational institutions, governmental agencies or nongovernmental organizations may apply for a Scientific Collecting Permit (SCP) to conduct research. The Department may authorize permitholders to pursue, capture, temporarily possess, temporarily injure, mark, attach to or surgically implant monitoring or recognition devices in, provide veterinary care to, and transport, mountain lions, or any part or product of a mountain lion.

Questions regarding the type of research being conducted as to whether a SCP permit is needed are encouraged to contact Dr. Lombardi. 

Approved Research Permitted Studies

There are currently seven active permitted research studies on mountain lions occurring throughout California examining topics including habitat connectivity, road ecology, genomics, energetics, response to human development, genetics and dietary analyses, and more.

More detailed information on each of these projects can be found here: Details of Current Permitted Studies

Prospective Researchers

Prospective researchers must submit a research proposal using a clear, bulleted format to facilitate the expeditious processing of the SCP request. Learn more!

  • Executive Summary;
  • Need Statement;
  • Time-oriented Objectives;
  • Study Area (include a map if applicable)
  • List of Personnel that details roles and responsibilities; and
  • Statement of how your project complies with the following criteria:
  • Contribute to the knowledge of natural wildlife ecosystems.
  • Minimize disruptions in the lives and movements of mountain lions and other wildlife, as well as impacts to mountain lion or other wildlife habitat, while maintaining the permitholder's research objectives.
  • Directly or indirectly support the sustainability and survival of mountain lion populations and healthy ecosystems.
  • Prevent the permanent injury or killing of any mountain lion.

If portions of any proposal contains intellectually confidential information (e.g., novel capture methodology), such information should be included in the appendix under a confidentiality statement heading that shall read: "The following information is a trade secret and should be exempt from public disclosure under the California Public Records Act pursuant to sections 6254(k) and 6276.44 of the Government Code." The proposal must still provide sufficient descriptive information, and simply reference this appendix when and where appropriate.

30-day Public Notices of Authorized Research Permits

The Department shall notify the public of the intent to issue a Scientific Collecting Permit (SCP) for mountain lion research 30 days prior to permit issuance, pursuant to Fish and Game Code Section 4810 . As of July 2023, CDFW has 1 pending SCP Permits for mountain lion research. Draft permit copies are available for review upon written request:

California Department of Fish and Wildlife Wildlife Branch – MOUNTAIN LION SCP 1812 Ninth Street Sacramento, CA 95811

To better inform management of of mountain lion conflict situations, the CDFW Human-Wildlife Conflicts Program and Mountain Lion Conservation Program work together to developed strategies to address the risk of mountain lions on the landscape using diverse tools and the best available science. Our highly-trained scientific and law enforcement staff address all cases of human conflicts with mountain lions. Training is a necessary part of an effective response to reported conflict. Communication, education, and outreach are important to enhancing the public’s understanding of potential conflict, conservation, and safe coexistence. For more information, visit the webpage and Keep Me Wildlife

For Human-Mountain Lion Conflict questions, please contact:

  • Victoria Monroe - Statewide Human-Wildlife Conflict Coordinator
  • Ryan Leahy - Statewide Human Wildlife Conflict Specialist

Human-wildlife conflicts are reported in urban and rural communities throughout California. As communities expand into wildland areas, increasingly reported interactions between people and mountain lions has resulted in increased calls for assistance from the public. Human population growth in the urban/wildland interface has manifested in small land parcels, many of which contain pets, small livestock, poultry, or exotic animals. These factors can lead to potential conflict due to increased attractants and competition for resources.

The Department seeks to avoid, where possible, mountain lion mortality resulting from the issuance of depredation permits. Further, staff strive to improve training, communication, transparency, and decision-making as they relate to managing human-wildlife conflicts involving mountain lions. It is recognized that each depredation incident may be unique. Property owners have legitimate concerns regarding mountain lion depredation. The Department understands these concerns, and is responsive to reported depredation pursuant to Fish and Game Code Section 4802 .

Since 2017, CDFW's approach has been deliberative, starting with a three-step policy implemented in the Santa Monica and Santa Ana Mountains. In February 2020, the policy was expanded to the larger proposed evolutionarily significant unit (ESU) contained in the petition to the Fish and Game Commission to list mountain lions under the California Endangered Species Act (CESA). Currently, staff use a deliberative stepwise process for responding to reported mountain lion incidents statewide.

A primary focus of the stepwise process is to prioritize approaches and actions that are non-lethal in accordance with Fish and Game Code Section 4801.5 , which provides that non-lethal measures shall be used when issuing depredation permits unless otherwise stated. Staff will provide property owners technical assistance, including:

  • Educational materials and resources regarding mountain lion behavior;
  • Potential reasonable preventative measures as recommended by the Department;
  • Site-specific options, including logistically and economically feasible measures to reduce the potential for attracting mountain lions;
  • Terms and conditions of a depredation permit, if applicable, and reporting requirements.

During the stepwise process, recommended measures may include, but not limited to: 1) removing the carcass and carcass parts of depredated animals; 2) install/repair/replace exclusion fencing; 3) implement more robust animal husbandry practices; 4) deploy temporary deterrent systems; 5) use livestock protection dogs; 6) hazing (e.g., use of bean bag shots), 7) capture, collar, and release measures to monitor movement and behavior.

Mountain Lion Depredation Statistics

A permittee is required to report take or no take upon expiration or fulfillment of the permit. CDFW maintains table summaries of the annual number of issued mountain lion depredation permits and the resultant take as reported to CDFW.

  • Annual Mountain Lion Depredation Summary Table (2011-2020) (PDF)
  • Annual Mountain Lion Depredation Summary Table (2001-2010) (PDF)
  • Annual Mountain Lion Depredation Summary Table (2020-2022) (PDF)
  • Data may be subject to change as new information becomes available.
  • These data represent the least number of permits issued to take a mountain lion and the least number of mountain lions taken under depredation permits in a given county in a given year. Temporal and/or geographic inconsistencies in reporting may make the data unreliable for identifying trends or making geographical comparisons without the application of a peer-approved scientific model that accounts for inconsistencies in reporting.
  • In some years, more lions were reported as taken than number of depredation permits issued. This may be due, but not limited, to inaccuracies in reporting. Additionally, multiple mountain lions could be taken on a single permit prior to 2013.
  • Depredation permits issued where the name of the county is not entered or legible are recorded as “Unknown County”.
  • Between 2016 and 2017, CDFW migrated all depredation permits to an electronic system.
  • Prior to 2018, the word “taken” is equivalent to “killed”. Since 2019, depredation permits which only authorize non-lethal methods under the new CDFW mountain lion depredation policy will be indicated separately in the table.

Threats to Public Safety

Mountain lions are typically solitary and elusive; often existing unseen and unheard. The Department receives hundreds of mountain lion sighting reports each year. Few sightings result in a mountain lion being identified as an imminent threat to public safety and killed under CDFW Wildlife Public Safety Guidelines. Most reports are resolved by providing species information and/or technical assistance. Cases where mountain lions threaten people are immediately addressed.

Mountain lion attacks on humans are rare. Statistically speaking a person is 1,000 times more likely to be struck be lightning than attacked by a mountain lion. Since 1890, there have been less than 50 verified mountain lion attacks on humans in California; of that, only six have been fatal. In most cases the person was lone when the attacked occurred.  List of known mountain lion attacks.

If a mountain lion is deemed a 'no harm no foul' animal and does not pose a threat, CDFW will work to encourage the animal back to its nearest suitable habitat. This may occur by monitoring and/or securing the local area to allow the animal to return on its own, actively hazing to deter it, o conducting a capture to relocate it. If a mountain lion displays unusually bold or aggressive behavior toward humans, the Department will not relocate the animal because of the risk it may pose to others. If the mountain lion is declared a public safety threat, CDFW and local law enforcement will work quickly to remove any threat in the most humane manner possible.

If a mountain lion is considered non-releasable (e.g., due to injury, disease, habituation), CDFW will work with permitted facilities and agency partners to try and located permanent placement of the animal. Most facilities, including wildlife sanctuaries have limited space or resources to accept large wild animals for exhibit.

The passage of the California Wildlife Protection Act of 1990 (Proposition 117) by California voters established that mountain lions are a "specially protected mammal" in California. It is unlawful to possess, transport, import or sell any mountain lion or part or product thereof (including taxidermy mounts). This status and other statues prohibit CDFW from developing hunting season or take limits for this species. The act established certain exemptions from that prohibition such as: mountain lions may only be harvested 1) if a depredation permit is issued to take an individual that has killed livestock or pets; 2) to preserve public safety; 3) to protect federally-listed bighorn sheep populations.

Specimen Possession / Taxidermy

No person may sell or possess any mountain lion, part or product thereof, unless s/he is in possession of a valid, nontransferable permit issued by the department. A permit shall be issued by CDFW with the following stipulations:

  • Any person who can demonstrate that the mountain lion, part or product thereof, was in his/her possession on or before June 6, 1990; or
  • A nonprofit museum or government-owned facility generally open to the public, or an educational institution, for display, exhibition, or storage, for a bona fide scientific or educational purpose as determined by the department.

Permits shall be made available to department staff for inspection, upon request, and may be revoked by the department for failure to comply with the terms of the permit, this section, or Section 4800 of the Fish and Game Code. Any person issued a permit or a tag from the department for a mountain lion, or part or product thereof, prior to January 1, 2014, shall not be required to obtain a new permit, provided s/he maintains and makes available his existing permit or tag upon request of a department employee.

  • Permit Application Requirements

Mountain lions typically pose little threat to humans, and generally avoid any human interaction. A person is one thousand times more likely to be struck by lightning than attacked by a mountain lion. People who live in mountain lion habitat can take precautions to reduce their risk of encountering a mountain lion.

Living With Mountain Lions

  • When recreating outside, avoid hiking, biking, or jogging alone. Do not hike, bike, or jog at dawn, dusk, or at night.
  • Deer-proof your property to avoid attracting a lion's main food source.
  • Remove dense vegetation from around the home to reduce hiding spaces.
  • Install outdoor lighting to make it difficult for mountain lions to approach unseen.
  • Secure livestock and outdoor large pets in sturdy, covered shelters at night.
  • Always remember - Mountain lions are wild animals and their behavior may be unpredictable (like any wildlife).

Mountain Lion Encounters

  • Stay alert on trails.  Keep pets leashed and walk with small children, don't let them run ahead. 
  • Never approach a mountain lion. Give them an escape route.
  • DO NOT RUN. Stay calm. Do not turn your back. 
  • Face the animal, make loud noise and try to look bigger. If with small children, put them on your shoulders. 
  • Do not crouch down or bend over.

Frequently Asked Questions

Related information.

  • Mountain Lion Depredation Statistics (Excel)
  • 'Keep Me Wild' Campaign
  • Human-Wildlife Conflicts Program
  • CDFW Memo: Amendment to Boundaries and Permit Requirements (PDF) - Effective 2/13/2020
  • Amendment to the Human / Wildlife Interactions in California: Mountain Lion Depredation, Public Safety, and Animal Welfare policy (2017) (PDF)
  • Human / Wildlife Interactions in California: Mountain Lion Depredation, Public Safety, and Animal Welfare policy (2013) (PDF)

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Cougars in Minnesota

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Cougar photograph

The cougar – sometimes referred to as a mountain lion or puma – was found throughout most of Minnesota prior to European settlement, though never in large numbers. Today, they are rarely seen but occasionally do appear.

While evidence might suggest the animal's prevalence is increasing, the number of verified cougar observations indicate that cougar occurrence in Minnesota is a result of transient animals from the Western Dakotas.

In addition, DNR annual scent-post and winter tracking surveys have recorded no evidence to suggest the possibility of a resident breeding population of cougars in Minnesota.

Although some cougar sightings in Minnesota are accurately identified, many observations from trail cameras and tracks turn out to be cases of mistaken identity. Bobcats, house cats, coyotes, wolves, fishers and light colored dogs have all been mistaken as cougars.

Because cougar observations are very rare, the DNR is interested in gathering information that can be verified by photos or physical evidence of a cougar sighting. Observations can be submitted to a local wildlife office or conservation officer.

Human encounters with cougars are extremely rare. Even in California, which has a population of more than 5,000 of the big cats, a person is 1,000 times more likely to be struck by lightning than attacked by a cougar. Most cougars will avoid confrontation. If an encounter should occur:

  • Face the cougar directly, raise your arms to make yourself appear larger and speak loudly and firmly. This behavior is in direct conflict with a cougar's tendency to hunt by stalking and attacking from ambush. Do not run, crouch or lay on the ground.
  • Do not shoot the animal , even if livestock or pets are threatened. Cougars are a protected species and may only be killed by a licensed peace officer or authorized permit holder.
  • Report the encounter or sighting to a conservation officer or local law enforcement authorities as soon as possible so evidence such as photographs, tracks, hair and scat can be located, identified, confirmed and documented.

Some cougars confirmed in Minnesota have had captive origins. Released or escaped pets often cannot be distinguished from wild animals until they are killed or captured and then only if they have obvious indications of captivity such as tattoos, de-clawing or tame behavior. Some cougars have not had any apparent evidence of being in captivity and may have been truly wild.

Since 2007, DNR has verified 14 sightings in Minnesota from trail camera photos of cougars, several cougars that were killed or found dead and other signs such as tracks. From dead cougars and biological samples collected, DNA evidence has linked these primarily to the populations in the western Dakotas.

U.S. Cougar Range

Cougar range in the U.S.

Click to enlarge

Several hundred miles separate Minnesota from the nearest known self-sustaining breeding population of cougars, estimated to number around 250, in the Black Hills area of South Dakota and, to lesser extent, the North Dakota Badlands. The only known population of cougars east of the Mississippi River is in Florida. Known as the Florida Panther, an estimated 90-100 wild animals continue to roam.

In the mid-2000s, a cougar that was fitted with a radio collar in the Black Hills was later located with telemetry equipment on the Roseau River Wildlife Management Area in northwestern Minnesota. This animal was there for about seven weeks before disappearing. It is possible that additional animals dispersing from the Black Hills or other western areas enter Minnesota.

Most animals confirmed in the Midwest during the past 15 years have been young males, the segment of a cougar population most likely to disperse in search of new territories. No females of suspected wild origin have been identified in Minnesota, and cougar tracks have not been recorded during annual scent-post and winter-tracking surveys. These facts suggest that the state does not have resident cougars.

It is most likely that transient animals (young male dispersers) are wandering into and through Minnesota. Once here, cougars are not restricted to territories by neighboring members of their species and can move freely.

While potential prey such as deer is abundant in Minnesota, dispersing cougars are not likely to stay in one area for long, instead continuing to search for suitable habitat with potential mates.

It is not unusual for people to mistake other animals for cougars. To identify a cougar, it is helpful to know that adult males can reach 200 pounds but most are much smaller and average about 150 pounds. Adult females usually weigh about 90 to 110 pounds.

The head appears small in relation to the body, which ranges in length from four to six feet overall. The body is tan except for dark face markings and the tip of the tail, which is nearly as long as the body.

Cougar sightings in Minnesota

Interactive map

There is no evidence that Minnesota has a self-sustaining, breeding population. And, because of their highly secretive nature and tendency to wander, it is nearly impossible to accurately determine how many individual cougars there might be in Minnesota.

During the past 30 years, DNR biologists doubt there has ever been more than a couple wanderers in the state at one time. Nevertheless, some sightings are confirmed, and cougar sighting confirmations in the midwest have increased in the recent past.

While uncertainty remains, most available information attributes the change to an increase in cougar populations in western states. Increasing populations produce more dispersing cats, particularly when those populations approach carrying capacity.

It is difficult to predict whether or when enough dispersing animals – both males and females – might settle in Minnesota and establish a small population.

Cougars subsist primarily on mammals, primarily ungulates such as deer, but also rabbit, squirrel, porcupine and other smaller animals.

When opportunity presents itself, cougars may go after farm livestock or pets. While the Minnesota DNR has received reports of cougars attacking livestock, there has not been an actual confirmed case of predation of livestock in Minnesota.

Report what you saw as soon as possible to the nearest area wildlife office or conservation officer . Note the date and time of the possible sighting, the location where the animal was seen and a description of what you saw.

If the opportunity presents itself, take a photograph of the animal or other potential evidence such as tracks. The DNR collects reports from citizens. Any physical evidence that could indicate the presence of a cougar will be investigated and proper steps taken to ensure public safety.

An encounter would be extremely rare in Minnesota. Cougars hunt by stalking and attacking from ambush and, if encountered should be, faced directly. Recommended actions include making yourself appear large by holding your arms above your head, waving a hat or jacket, talking loudly and firmly and throwing rocks or sticks at the animal to chase it away.

If actually attacked, hit the animal in the face and head with anything handy. Don't run, crouch or lay down. Try and stay above the animal and give the animal a clear escape route.

No. Cougars are protected by state law. Only public safety officials are authorized to kill a cougar that proves to be an imminent threat to humans.

If there is a proven public safety concern, DNR or other licensed peace officers can take a cougar to protect public safety. State law does not allow killing a cougar to protect livestock or pets.

Should you become concerned with a cougar in your vicinity, contact a DNR conservation officer or local law enforcement authorities to evaluate the situation. Local law enforcement has the responsibility for public safety. DNR enforcement and wildlife personnel will work with local law enforcement to quickly evaluate and resolve problems.

Relocating problem animals usually is impractical because the same problems could occur at new release sites. Due to the transient nature of cougars in Minnesota, it is extremely difficult to capture a cougar based on an individual incident.

Deporting them to western states, where cougar populations already exist, is often prohibitively expensive and can also result in territorial struggles and the death of the released animal.

More information

  • Rare species guide
  • Cougar treks 2,000 miles east

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mountain lion travel range

How California is helping its mountain lion population

H ighway 101 in California is closed starting this week as workers start construction on a wildlife corridor that will be “the world’s largest wildlife crossing,” according to CNN .

The 101 wildlife crossing project started because a lone mountain lion, known as P-22, would often roam Griffith Park, but was unable to cross the 101 to find a mate, CNN reported back in 2022.

Construction of the 101 wildlife crossing

CNN noted that Hollywood celebrities donated money to support the 101 wildlife crossing officially known as the Wallis Annenberg Wildlife Overpass.

The Wallis Annenberg Wildlife Overpass will connect mountain lion populations that are separated by the 101 highway, according to the National Park Service .

The overpass will help resolve scientists’ fears about inbreeding among mountain lions in Southern California, the National Park Service said.

Expected to be completed in 2025, the wildlife crossing will be the biggest one in the United States and will have natural features such as native plants to help it blend into the natural environment, per CNN .

What’s a wildlife crossing?

The U.S. Fish and Wildlife Service explains that wildlife corridors are paths that various species travel in response to weather events, seasonal changes or life cycle milestones.

For example, the National Elk Refugee in Wyoming is what elk in the area use to migrate in the winter.

Human development, including highways, can threaten natural wildlife corridors. That explains why many communities have built bridges, tunnels and other wildlife crossings to make up for changes to natural corridors.

Without them, many species can’t find food or mates, according to National Geographic .

National Geographic explains the several forms a wildlife crossing can take:

  • Wildlife bridges: First developed in France in the 1950s, wildlife bridges are covered in natural vegetation that help create a more inviting environment for animals. The longest wildlife bridge is in Denmark that measures 0.5 miles long.
  • Ecoducts: Ecoducts include a tunnel that runs underneath a road originally built for toads in Davis, California. It has not been very popular among the toads.
  • Wildlife crossings: This umbrella term that refers to bridges or tunnels that allow animals to safely cross highways and have been shown to decrease animal-related car accidents.

An overview of the Wallis Annenberg Wildlife Crossing, which will eventually be built over the 101 freeway, Tuesday, Sept. 20, 2022, in Agoura Hills, Calif. The construction for the 101 wildlife crossing started because a lone mountain lion known as P-22 would often roam Griffith Park, but was unable to cross the 101 to find a mate.

The Pet Savvy

107 Animals That Start With M

By Eddie Chevrel

Updated on June 3, 2022

animals that start with the letter M

What animals start with the letter M? We’ve gathered them all here for you to have them in one place – from the majestic manta ray to the annoying moth, right through to the mink and the mole.

So without further ado, here is the list of animals that start with the letter M.

Animals Starting With The Letter M – The Complete List

Macaroni Penguin

Mackenzie Wolf

Madagascar Hissing Cockroach

Madagascar Tree Boa

Magellanic Penguin

  • Mahi Mahi (Dolphin Fish)

Malayan Civet

Malayan Tiger

Maltese Shih Tzu

Mamushi Snake

Man of War Jellyfish

Manchester Terrier

Mandarin Rat Snake

Mangrove Snake

Mantella Frog

Marabou Stork

Marine Iguana

Marine Toad

Masked Angelfish

Masked Palm Civet

Megamouth Shark

Mekong Giant Catfish

Mexican Alligator Lizard

Mexican Black Kingsnake

  • Mexican Eagle (Northern crested caracara)

Mexican Free-Tailed Bat

Mexican Mole Lizard

Mexican Red Knee Tarantula

Midget Faded Rattlesnake

Mini Labradoodle

Miniature Bull Terrier

Miniature Husky

Miniature Pinscher

Minke Whale

Mississippi Kite

Moccasin Snake

Mojave Rattlesnake

Mole Cricket

Monarch Butterfly

Monitor Lizard

Monocled Cobra

  • Monte Iberia Eleuth

Moon Jellyfish

Moscow Watchdog

Mountain Bluebird

Mountain Cur

Mountain Feist

Mountain Gorilla

Mountain Lion

Mourning Dove

Mulga Snake

Mussurana Snake

The Macaroni Penguin is a species of penguin that lives in the Subantarctic to the Antarctic Peninsula. They are sometimes known as “Penguin Mike” because they make a specific sound when puffing out their chests. This is thought to be a call for mates, but it’s not clear why only males do this.

The macaw is a parrot that comes in many different colors and sizes. The largest macaws are at least two feet tall, while the smallest is only 10 inches long. Macaws live in South America and Central America, where they eat fruits, nuts, seeds, flowers, and leaves from trees such as Brazil nut trees and acai palm trees.

Mackenzie Wolf is a subspecies of Grey Wolf that lives in the northwest area of North America. It is named after explorer Alexander Mackenzie and has been known to attack humans when provoked or looking for food.

The Madagascar Hissing Cockroach is a species of cockroach that can grow up to 3 inches long! They live in Madagascar and eat plants and other insects.

The Madagascar Tree Boa is a large snake found only on Madagascar Island, where it lives in trees and eats small mammals such as rats or mice. These snakes can grow up to 10 feet long!

This penguin can be found in South America and the Falkland Islands. They swim in cold water and have blackheads and white bodies. Their diet consists of fish, squid, crabs and krill.

Maggots are fly larvae that feed on dead animals or decaying matter; they are sometimes used as fishing bait because they attract fish but don’t kill them (like conventional bait).

Magpies are members of the Corvidae family and are native to Europe and Asia. They have black and white feathers with a yellow beak and feet. They eat insects, small mammals, seeds, berries and other fruit. They have an average lifespan of 5 years in the wild but up to 15 years in captivity.

Mahi Mahi is a type of fish that lives in tropical oceans worldwide with an average length of 1-2 meters long! It can be found mostly near coral reefs or rocky bottoms but also in open water if there are enough food sources nearby like algae or plankton, for instance!

The Maine Coon is a breed of domestic cat. It was developed in New England and is one of the largest domestic breeds, known for its particularly long body and strong muscles. The cat has a distinctively long tail and large paws.

The Malayan civet is a viverrid native to Southeast Asia. It is found in Indonesia, Malaysia, Myanmar, Singapore, Thailand and Vietnam. The civets are solitary animals that feed on insects and small vertebrates such as birds and rodents. They are also known to prey upon lizards and snakes.

The Malayan tiger is an endangered subspecies of tigers found in the forests of Indonesia and Malaysia. Once common throughout Southeast Asia, their numbers have declined due to hunting and habitat loss. Today, fewer than 500 tigers are left in the wild out of an estimated total population of 3200 tigers worldwide!

Mallards are members of the duck family Anatidae found in Europe and North America and parts of Asia and Africa. They are often kept as pets because they are easy to care for, but they may become aggressive during mating season when they feel threatened by other males or humans who get too close while trying to mate with females!

A breed of dog that has been around since ancient times. They are small but very intelligent and easy to train. They are also known for their love of human affection, making great companions for people who like pets that can cuddle with them!

A cross between a Maltese and a Shih Tzu dog, this mix was created by breeders in order to get the best qualities from both breeds-the intelligence of the Maltese combined with the playful nature of the Shih Tzu.

Breeders created a cross between a Maltese and a Toy Poodle, this mix to get the best qualities from both breeds-the intelligence of the Maltese combined with the playful nature of the Toy Poodle.

The mamushi snake can be found throughout Japan and China and is one of the most venomous snakes globally. It has an aggressive temperament and will attack humans who get too close to it.

Also known as blue jellyfish, this species is found all over the world’s oceans. Its bright blue coloration makes it easy to spot in the water, but its sting is not deadly to humans-just excruciating.

Manatees are large marine mammals that live off Florida’s coast in shallow waters. They have rounded bodies with short heads that are sometimes called “sea cows.” Their skin is grayish brown and covered with fine hairs called “whiskers,” which help them navigate their murky habitat.

This small dog breed was developed in Manchester, England, during the nineteenth century. They have compact bodies with long legs, which make them great jumpers! They come in many different colors, including black and white.

The Mandarin Rat Snake is a non-venomous snake that lives mainly in China but can also be found in Taiwan and Singapore. It has a brownish back and orange belly with black spots on it.

The mandrill is a colorful baboon found in Africa. They have large, hairless heads with bright orange or red cheeks and blue buttocks. They also have large nasal cavities used to make loud calls when hunting for food or warning others of danger.

The maned wolf is native to South America. Their name comes from the long reddish-brown hair on their backs that resembles a mane. They are omnivores and eat both plants and animals such as rodents, birds, insects, and reptiles. They also scavenge carcasses when they can find them, which helps them get nutrients from meat without killing anything themselves!

The Mangrove Snake is a non-venomous reptile found in coastal regions worldwide. They have large triangular heads with sharp teeth designed for eating fish, which makes them great hunters! They also have an excellent vision that allows them to see their prey far away-even at night!

The Manta Ray is one of the largest ray species in the world, growing up to 20 feet wide! These creatures can be found swimming in warm tropical waters all over the globe; however, they are most commonly seen near coral reefs, where they feed on plankton, small fish, and crustaceans. Their bodies are covered in tiny tooth-like scales called denticles (made out of calcium carbonate) which protect them from predators.

This frog has been known for its medicinal properties for thousands of years. It contains an alkaloid called D-tubocurarine, which helps relax muscles during surgery or childbirth by blocking electrical signals from nerves to muscles. There are currently only five species left in the world!

The marabou stork is a large bird that lives in Africa and Asia. It can be found in the savanna or open country near water. The name “marabou” comes from the Arabic word for “bald.”

The marble fox is a small fox living in northern China and Mongolia. It has white fur on its body and black fur around its eyes. The name “marble” comes from the appearance of this animal’s fur when it is wet – it looks like marble!

The marine iguana is an iguana that lives in the Galapagos Islands off South America. These animals feed on seaweed, algae and other sea plants underwater. They have special adaptations for living underwater, such as webbed feet and nostrils on top of their heads so they can breathe while submerged!

The marine toad has a large flat head with bulging eyes and wide nostrils. Its skin is reddish-brown and covered in warts. It has webbed feet and long, sticky fingers that help it catch prey in water.

The markhor is a wild goat found in Afghanistan. It has long legs and horns that curve backward from its head like scimitars. The male markhor’s horns can grow up to 6 feet long!

The marmoset is native to South America and gets its name from its resemblance to monkeys of the same name found in Africa. They are very small animals-usually only six inches tall-and have large heads with rounded foreheads, and large eyes set close together on either side of their face. They are also called “little monkeys” because they act like monkeys sometimes (like when they eat their poop).

Marmots are rodents that live in cold climates. They are large and covered in fur, making them look like big fluff balls! Their bodies are a light brown with white patches on their faces and bellies. Marmots hibernate during the winter months when it gets cold outside.

The marsh frog is one of several species of amphibians that can be found around the world today. This amphibian has long legs, which help it hop around its habitat much more easily than other frogs do! It is usually brown or greenish-brown in color with some darker brown spots on its back legs (which appear like stripes). It can grow up to eight inches long-much larger than most other frogs!

The masked angelfish lives in coral reefs around Southeast Asia and Australia, where it feeds on coral polyps by eating their tentacles before destroying their bodies with its sharp teeth! The fish gets its name from its bright colors: yellowfins with orange stripes.

The masked palm civet is a nocturnal mammal native to Southeast Asia. It is also known as the banded palm civet or Zibetto palm civet, which refers to its striped coat. It has large eyes and small rounded ears close to its head.

Massasaugas are small snakes with large triangular heads and glossy scales. They tend to be brown or black, with yellowish bellies or throats. Massasaugas have been found throughout North America, including southern Canada and the United States.

Matadors are large dogs initially bred in Spain as herders and guard dogs for cattle ranches. Still, they have since been used as guard dogs for other animals such as sheep and goats and pets owned by people living in rural areas where there may be no fences around their homes or yards (such as farms). They can weigh up to 175 pounds!

Maltese Eagle Dog

Maltese Eagle Dogs is a small-sized dog breed. They have a long, silky coat in white and black or white and gray. These dogs are known for their hunting ability and their loyalty to their owners.

A Mastiff is a dog breed. It is one of the giant breeds of dogs in the world. The Mastiff is an ancient breed that has been around for centuries. The Mastiff is known to be gentle and loving with its owners, but it can be aggressive towards other animals and people.

The mayfly is an insect that only lives for a few hours after it emerges from its watery egg case. The insect usually travels around looking for food before it dies at nightfall. Mayflies are found all over the world, except in Antarctica.

The meagle is a small mammal found in Australia that has a very long neck and legs that help it reach high branches where it eats leaves and fruit from trees. Meagles are also known as wallabies because they climb over walls to escape predators like dingos or foxes that try to catch them for food or fun! Humans have hunted this species for thousands of years because they have fur on their backs which makes nice warm clothing during winter months when temperatures drop below freezing point!

A Mealybug is a small insect that feeds on plants and fruits. Its name comes from its white mealy substance, which it produces when it feels threatened by predators such as ants or spiders.

Meerkats are African mammals that live in groups called “mobs.” They are omnivores and feed on insects, small reptiles, amphibians and rodents. Their main predators are jackals, snakes and large birds.

Megalodon is an extinct shark species that lived between 23 and 2.6 million years ago. It was one of the giant sharks ever and probably reached a maximum length of 17 m (55 ft). Its teeth were up to 7 inches long! The megalodon became extinct about 1.5 million years ago after a decline starting around 1 million years ago.

The megamouth shark (Megachasma pelagios) is a rare lamniform shark that inhabits deep-water environments worldwide. It is characterized by a large head and a broad mouth lined with numerous rows of tiny teeth.

The Mekong giant catfish, Pangasianodon gigas, is the largest species of freshwater fish in the world, reaching up to 10 feet in length and weighing up to 300 pounds. It can be found in Southeast Asia and is listed as vulnerable by the IUCN due to overfishing and habitat loss.

The Mexican alligator lizard (Elgaria coerulea) is an endangered species that lives in Mexico and the United States. It is commonly confused with its cousin, the common garter snake because they look almost identical but have different habitats.

The Mexican black kingsnake (Lampropeltis getula nigrita) subspecies of the common kingsnake in Mexico and southern Arizona. It can grow up to six feet long and has dark brown blotches on its back. It’s often mistaken for a venomous coral snake because of its similar coloring.

Mexican Eagle

This bird is a large raptor with a yellowish-brown body and black wings. It is also called the Northern Crested Caracara. It has a crest of feathers on its head and a black mask around its eyes. It lives in Mexico and Central America, and hunts for food at dusk or dawn.

The Mexican free-tailed bat is one of the most common types of bat in North America. It lives throughout Mexico and parts of the United States, where it roosts in caves during the day before flying out at night to hunt for insects.

The Mexican mole lizard is native to Mexico and Central America. It has a long body that curves upward at the end like an S, so it looks like it has two tails! It eats insects, small reptiles, and worms that live underground-hence its name!

The Mexican red knee tarantula is a type of large spider that is native to Central America. These spiders are characterized by their black bodies and red legs. They are popular in the pet trade.

A snake found in sub-Saharan Africa that can grow up to ten feet long! It gets its name because its movement resembles that of a snake moving through water (mamba = mambo).

The muntjac is a small deer species that lives in Asia. They are very good at eating bamboo shoots, and they can jump high! They are like rabbits in that way, but they’re not rabbits-they’re just related to them.

The Muskox is an Arctic animal that lives in northern Canada and Greenland. It has large horns on the top of its head which protect it from predators such as wolves or polar bears. Muskoxen are also very fast runners, so they can outrun most predators if need be!

The muskrat is a small rodent found throughout North America. They often live in wetlands or swamps and are known for building dams with mud and grasses to create ponds where they can swim around all day long without getting wet (or cold).

The Midget Faded Rattlesnake is a small snake species found in the Southern United States. It has a brown body with black stripes running down its back and a white underbelly. The Midget Faded Rattlesnake can be identified by its faded coloration and small size.

Miki is a Japanese word that means “a small amount.” Miki is Japanese short-legged dogs with large heads and eyes, making them look like they’re always surprised. They’re also very intelligent and affectionate, making them great pets for families with children.

When threatened or agitated, the milk snake gets its name from the milky-colored liquid it secretes. This nonpoisonous snake can grow up to five feet long, though most will only reach three feet. They live in wooded areas throughout the U.S., Canada, Mexico, Central America, and South America.

This fish is native to the Indo-Pacific region, including the Red Sea and East Africa. It can grow up to five feet long and weigh up to 100 pounds. Its flesh has a mild flavor and firm texture, making it popular in many cuisines throughout Asia and Africa.

This creature has been around since before dinosaurs roamed the Earth; fossils dating back 400 million years have been found with millipedes inside them! Millipedes are typically between one and two inches long (though some species grow much larger). Still, they possess an incredible ability: they can produce hydrogen cyanide as a defense mechanism against predators.

The Mini Labradoodle is a cross between the Labrador Retriever and the Poodle. They have many of the same characteristics as their larger counterparts, but they are generally more energetic and playful.

The Miniature Bull Terrier is a smaller version of the traditional English Bulldog. They are still characterized by their wrinkled faces and short muzzles, though they tend to be more active and energetic than their larger counterparts.

The Miniature Husky has been bred down from the Siberian Husky to be much smaller in size than its parent breed. It has the same characteristics as a full-sized Husky, but it tends to be more playful and affectionate than other breeds in its family tree.

The Miniature Pinscher is one of several breeds bred down from larger dogs like German Shepherds or Rottweilers to be much smaller in size than traditional versions of these breeds would normally be expected to reach maturity. They are known for being brilliant dogs who can be trained easily with patience and consistency on behalf of their owners.

Mink are small, carnivorous mammals that are closely related to weasels. They are found in North America, Europe, and Asia. The mink’s fur is highly prized for its softness and warmth, and many people hunt them for their pelts.

The minke whale is a type of baleen whale that lives in cold waters worldwide. It can grow up to 20 feet long and weigh up to 8 tons. It has a dark gray body with white patches on its underside and a white band around its head. Minke whales feed on krill and small fish, using their baleen plates to strain these prey items from water passing through them as they swim through it at high speeds (upwards of 30 miles per hour).

The Mississippi kite is a large bird of prey in the family Accipitridae. It is native to North America but can also be found throughout Central America and South America. It is named after the Mississippi River, where Europeans first discovered it. Its preferred habitat is open country with scattered trees, such as riverbanks and fields with scattered bushes or forest edges.

The moccasin snake is a venomous pitviper species native to the southeastern United States from Texas through Georgia and Florida. Its coloration varies from yellowish-green to brownish-black, with lighter coloration on its belly region.

The Mojave rattlesnake is a venomous reptile found in southwestern United States desert regions. Its venom contains a neurotoxin that can cause severe pain, swelling, and paralysis.

The Mola mola is not a fish but a type of ray. It lives in tropical oceans worldwide and has huge scales that help it float on top of the water without sinking. The fish can grow up to 10 feet long and is often referred to as the Ocean Sunfish.

A mole is a small burrowing mammal with pink or gray fur that lives underground. Moles have tiny eyes and ears covered by fur, so they rely on their noses to find food and avoid dangers like predators or traps set by humans who want to kill them for their fur.

The mole cricket is a small insect that lives in tunnels underground. It has large, powerful front legs and a long, thin body that it uses to dig. It eats plants and small insects.

The mole snake is a burrowing snake that lives in holes underground. It has small eyes and a thick body and is covered in scales. Like other burrowing animals like moles and gophers, the mole snake can excavate large amounts of dirt with its strong front legs.

Mollies are brightly colored fish with long fins to swim gracefully through the water. They are among the most popular aquarium fish because they come in so many different colors. Mollies eat algae and plankton, which they filter out of the water using their gills. They are also known to eat smaller fish if they get hungry enough!

Monarchs are one of the most recognizable butterflies in North America because their wings have orange and black stripes (which makes them look like monarchs). They eat only milkweed plants because they contain chemicals called cardenolides which make them poisonous to predators like birds.

Mongooses are mammals that live in Africa and Asia. They have long bodies, short legs, and small ears. The mongoose has a very strong sense of smell which helps it hunt for food. Mongooses also have sharp teeth, which they use to kill their prey.

A mongrel is a dog that has been bred from different breeds of dogs. Mongrels can be mixed with many kinds of dogs, which gives them the ability to adapt to different environments.

The monitor lizard is one of the giant lizards in the world. It can grow up to 3 meters long and weigh over 30 kilograms. The monitor lizard has sharp claws on its feet to catch prey such as insects and small animals like mice or rats.

Monkeys live in tropical rainforests worldwide, with two species living in Africa and South America being found nowhere else on Earth! Monkeys live in large groups called troops, usually consisting of about 20-50 individuals each; these groups can be made up of both males and females but will always include at least one male monkey called an alpha male.

The monkfish is a type of anglerfish. It has a long, slim body that can reach up to 2 m in length. This fish lives at depths of up to 1,000 m and catches its prey using its large mouth and sharp teeth.

The monocled cobra is found in South Asia and Southeast Asia. It has a triangular head with distinctive eye markings, plus a yellowish or orange belly and a black body with light-colored bands on the back. It is thought that this snake may be venomous, but it’s not yet known how dangerous it is to humans.

Monte Iberia Sleuth

This rodent weighs around 1 kg (2 lb) when fully grown and displays unusual physical characteristics such as large ears and tail and an unusually short snout for its size, which resembles an anteater’s snout.

The moon jellyfish belongs to a group of jellyfish called Aurelia aurita, also known as common jellyfish or lunar jellies because they resembled the moon when seen from space! They’re found worldwide in oceans, both warm and cold.

Moorhens are members of the rail family and are known for their ability to fly. Their wings are made of almost entirely black feathers, which makes them difficult to see when they’re flying. They live in freshwater habitats like lakes and ponds.

Moose are large members of the deer family that live in northern forests and tundras. They have a thick coat of fur that keeps them warm during cold weather, and their antlers can weigh up to 90 pounds!

Moray eels are carnivores that live on coral reefs in tropical waters worldwide. They’re easily recognizable by their large jaws filled with razor-sharp teeth-which they use to catch prey such as fish and crustaceans!

Morkies are hybrid dogs bred from Yorkshire Terriers and Maltese dogs. They’re popular pets because they’re small but sturdy enough to be good watchdogs (they bark loudly at strangers), but also affectionate and friendly towards people they know well enough not to bite them!

Moscow Watchdogs are a breed of dog developed by Russian scientists after World War II to guard nuclear weapons sites within Moscow city limits against intruders.

Mosquitoes are small, flying insects that live in wet areas. They have a long proboscis which they use to suck blood. Their bite can be painful and cause an itchy red bump.

Moths are winged insects that belong to the Lepidoptera order. Most moths are nocturnal or crepuscular, meaning they feed during the night or at dawn and dusk.

Mountain bluebirds are small birds that live in high altitudes and prefer open areas like meadows and fields with brushy undergrowth. They eat seeds from plants like grasses, sedges, and rushes and berries from shrubs such as wild rose bushes or currants.

A mountain cur is a type of dog commonly used for hunting in mountainous regions where other breeds would tire easily due to their short legs and thick coats. They come in several different colors, including black/tan (also known as brindle), blue merle (black with white patches), red merle (red with white patches), sable (black with brown markings), and yellow.

This dog is a cross between a terrier and a dachshund, known for its aggressive attitude. It was originally bred to be a hunting dog, but its small size made it ideal as a companion dog instead.

The mountain gorilla is an endangered species of gorilla native to the Virungas, which are five volcanoes located on the borders of Rwanda, Uganda, and Congo. These gorillas are known for their gentle nature and ability to use tools for foraging food. They are one of the few primates who have been observed using tools in their natural environment-they will tear the bark off of trees to eat insects underneath them or use sticks as tools for digging up roots.

This cat lives in mountain ranges across North America and is a relative of both cougars and bobcats. Mountain lions have been known to kill humans, though they typically avoid people and are more likely to try to avoid them than attack them.

This bird is found all over North America, though it prefers open areas with tall grasses or shrubs where it can build its nest-which it does communally with other mourning doves in large colonies of up to 50 nests!

Mice are rodents that live everywhere, from deserts to forests-though some species prefer specific climates. They tend to travel by running rather than walking to move quickly through their environments.

The mud snake is a non-venomous snake native to the southeastern United States. The mud snake has an olive, brown, or black color with a yellowish belly. It is typically about 36 inches long.

The Mudi is a Hungarian herding dog that looks similar to an Airedale Terrier. They have prick ears and a medium-length coat that can be either solid white or white with patches of red or tan on their backs and legs.

The mudpuppy is a North American amphibian that lives in swamps and marshes across the southern United States. The mudpuppy has five toes on each foot, which helps it burrow into the mud when threatened by predators such as snakes and birds of prey like hawks and owls.

A mule is the offspring of a donkey stallion and horse mare, both of which are members of different species (the horse belongs to the genus Equus while donkeys belong to the genus Asinus). Mules are sterile but can reproduce through artificial insemination if they mate with another mule or donkey.

Mulga is the name of a minor, venomous snake that lives in Australia. It’s also the name of one of my favorite snakes because it’s so cute! And cuddly. And I like to snuggle with them.

The mussurana is a non-venomous snake that originates from South America. It has a long body and can grow up to 7 feet (2 meters) in length. The mussurana has a flat, triangular head and small, dark eyes. It can be found near water in Brazil, Colombia and Venezuela.

The myna bird is native to India but has been introduced to many other parts, including Australia, Indonesia and Hawaii. The myna bird is known for its ability to learn human speech and imitate sounds such as doorbells or truck engines.

Mojarra is a freshwater fish found in the Amazon River basin. It is a popular food fish, and it is also used as bait for other species. Mojarra thrives in brackish waters with low oxygen levels; however, their populations often decline when introduced into areas with higher oxygen levels.

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About Eddie Chevrel

Eddie Chevrel is an animal journalist and the founder of ThePetSavvy. He's very passionate about exotic pets and spends most of his free time doing research, meeting, and interviewing people working with animals. Learn more about The Pet Savvy's Editorial Process .

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Last updated: May 4, 2018

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Contact info, mailing address:.

1000 US Hwy 36 Estes Park, CO 80517

970 586-1206 The Information Office is open year-round: 8:00 a.m. - 4:00 p.m. daily in summer; 8:00 a.m. - 4:00 p.m. Mondays - Fridays and 8:00 a.m. - 12:00 p.m. Saturdays - Sundays in winter. Recorded Trail Ridge Road status: (970) 586-1222.

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Moscow parks – leisure, nature and historical

Moscow is the most green megapolis in the world. There are over a hundred parks and green spaces like gardens, squares and boulevards. You will definitely bump into a few of them wherever you go. Whether you are interested in memorial, historic parks, parks of wildlife or you just want to have a calm break from the speedy city life – city parks have something special for everyone.

Moscow leisure parks

The leisure Moscow parks are undoubtedly the most popular and famous with the locals and travelers. Today such parks provide a great number of exciting entertainments for Muscovites and city guests, adults and children.

The Gorky Park

The Gorky Park

Gorky Park opened in 1928 and was the first holiday park in the Soviet Union with playgrounds, a sports stadium, exhibition halls and attractions for kids. Today it has a fresh, vibrant appearance. The park features bike rental stations, a comfortable business area with Wi-Fi, an outdoor movie theatre and a greenhouse where you can buy fresh greens such as basil and lettuce. To contemplate the sky and the stars, go to the observatory and look through the telescope while listening to fascinating stories from astronomers. Enjoy many sports in the park: volleyball, handball, football or a peaceful jog around the beautiful surroundings.

Zaryadye Park

Zaryadye Park

Opened in September 2017, Zaryadye is the youngest on our list. Located just a few minutes away from Red Square, it includes various activities like the floating bridge with its thin V-form extension, an ice cave, also concert hall and an amphitheater. The entire territory of the park was divided into four zones of Russia: forest, steppe, tundra, and the floodplains.

Neskuchny Sad

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By walking along the Moskva River’s bank from the Gorky Park towards Vorobievy Gory (Sparrow Hills) you’ll reach Neskuchny Sad («Not Boring» garden), a wonderful place in the Moscow center, one of Moscow’s oldest parks, charming slice of wildlife. The park mostly consists of pristine forest, dotted with old summer pavilions, ponds and quaint little stone bridges. There are a lot of opportunities for different activities lots of children playground, a ping-pong and chess clubs, football fields and tennis courts, horse riding, tree climbing and having rest in one of the nice cafes.

Museon Park

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Hermitage Garden

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Hermitage Garden has always been known as an amusement, entertainment center with theatres, shows, cafes, summer pavilions, pergolas since 1830. Shalyapin, Sobinov, Nezhdanova – great Russian opera singers starred on the garden stage. Famous Russian composer Rakhmaninov conducted the orchestra. Sara Bernar, Maria Yermolova, outstanding actresses, played in the open air performances. Tolstoy and Lenin had a stroll in the garden. So lots of celebrities from different epoques liked it a lot and spent their time in Hermitage Garden. You can find here three theatres in the garden: Hermitage, Sphere and New Opera. During winter an ice rink works here and in summer a musical stage is assembled to host jazz and brass band festivals.

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In Sokolniki Park visitors can play billiard, chess or draughts, table tennis, as well as go cycling, roller blading and swimming in the summer and ice skating or skiing in winter. Each season is highlighted by special memorable and bright events, for example, Summer Jazz Festival or Baby Fest (for future mums), open air beach disco parties, Ice Cream Day, International Clown Festival and many other shows and exhibitions. The park has an observatory, kids center and a co-working zone with free Wi-Fi which is really nice for spending high quality work time there.

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Moscow nature parks

The nature parks are national reserves with the amazing forestry and incredible variety of animals and plants there. The breath of wildlife and the chance to be closer to the virgin nature excites both children and grownups. Hundreds of different species of animals can be found in Moscow nature parks. The richest woodlands with old and even ancient trees, like a 200 years pine-tree in the Elk Island National Nature Park, are the point of passionate interest for visitors.

Aptekarsky Ogorod

Aptekarsky Ogorod (Apothecary Garden)

Aptekarsky Ogorod (Apothecary Garden)

Aptekarsky Ogorod (Apothecary Garden) is one of the oldest gardens in Moscow. It was founded in the XVIII century by Peter the First (great Russian emperor). A larch that he planted himself still grows in the garden, so it’s more than 250 years old. At the time of its foundation, it was a garden with herbs and medicinal plants and was used as an educational center for doctors. Today there are the orangery with its tropical palms, the carp pond, and the immense trees that dot the landscape and turn wonderful golden shades in the autumn. Several restaurants and cafes work here making it a very nice spot for relaxation no matter what season it is. You can book a special tour or join the guided excursion group.

Losiny Ostrov

Losiny Ostrov (Elk Island Park)

Losiny Ostrov (Elk Island Park)

Losiny Ostrov (Elk Island Park) is located at the north of Moscow. It covers 22 km from the west to the east and 10 km from the north to the south and it’s one of the most beautiful national parks in Moscow. Two rivers, Yausa and Pechorka begin here. You can find lots of fields, ponds, meadows, streams in the park as well as elks. Here you can enjoy guided tours, available in English. You can choose a guided tour about flora and fauna of the area, you’ll learn why elks are there, which animals are their neighbors. Or enjoy another excursion, which is totally devoted to historical past of ancient tribes once lived there, you’ll know about old Russian mythology, rituals and traditions.

Serebyany Bor

mountain lion travel range

Serebyany Bor (Silver Forest)

Serebyany Bor (Silver Forest) is a famous pine forest in the west of Moscow. The park has 230 forms of plant life, and is also home to watersports complex, providing a lot of activities for visitors. The layout of Serebryany Bor is unusual, as it is located on an artificial island between a meander in the Moscow River and a channel. There is an artificial lake, the Deep Gulf and picturesque Bezdonnoe (Bottomless) Lake in the depths of the forest. Serebryany Bor’s beaches are the cleanest in the city and very popular among Muscovites. On weekends it is difficult to find a free spot here, especially because a whole range of services are offered to visitors, from simple deckchairs to catamaran and yacht rides. Driving is prohibited on the territory of the island so be ready to use trolleybus to reach the entrance.

Botanichesky Sad

Greenhouse of Botanical Garden

Greenhouse of Botanical Garden

Main Botanical Garden of The Russian Academy of Sciences is the largest and most famous is Moscow. The garden is a real museum of nature with a very rich (more than 18000 types) collection of plants. The park was founded in 1945 at the place of the 17th century Apothecaries’ Gardens. The garden’s collection is turned into botanical expositions, made with use of modern receptions of landscape architecture. Here you can see a tree nursery, a shadow garden, hothouse complex, collection of flowers, a rosary, exposition of coastal plants, garden of continuous blossoming, Japanese garden and expositions of cultural plants and natural flora plants. The biggest part of Garden is the Tree nursery occupying the space of 75 hectares. About 2 thousand wood plants grow here. Another big exposition of the Garden is nature Flora, divided into six botanic-geographical collections: European part of Russia, Caucuses, Central Asia, Siberia and Far East. Pride of the Main Botanical Garden is the collection of tropical, coastal and water plants, which is considered as the best in Europe. The Japanese garden, a great model of Japanese landscape gardening art has a 13-level stone pagoda of the 18th century, stone Japanese lamps, ponds, falls and streams, tea lodges and more than 100 species of the most character Japan plants. It is especially decorative in spring, during Oriental cherry blossoming and in fall, when foliage blazes in crimson colors.

Moscow historic parks

Historic nature parks and estates once were the mansions of the Moscow aristocracy. At that far times the estates were outside the Moscow city limits, but after the city expansion and urbanization, they became easily accessible.

Kolomenskoye

Kolomenskoe

Kolomenskoye Museum and Park

The chief attraction of the park is undoubtedly the stone Church of the Ascension of the Lord. It was constructed in 1532 by order of Tsar Vasily III to commemorate the birth of his son and heir, Ivan the Terrible. But there is a lot more to see in the park: the pretty Church of the Icon of Our Lady of Kazan – with its bright azure domes and plenty of gold. Further into the park there is a charming Church of the Beheading of St John the Baptist, built by Ivan the Terrible to mark his coronation.

Kuskovo

Kuskovo Park

Kuskovo Park is one of the oldest country estates in Moscow. It was given to General Sheremetev by Peter the Great in 1715, but was left to fall into neglect before being plundered by Napoleon’s troops in 1812. Nowadays the estate has been restored to its former glory and is a good example of Russian 18th Century imperial architecture. The palace is a fine and rare example of wooden neoclassicism. It was completed in 1775, and the rich interiors remain unchanged since 1779. It includes a room hung with exclusive exquisite Flemish tapestries, an abundance of silk wallpaper and an impressive collection of 18th century European and Russian paintings. The palace looks onto the lake, which is surrounded by smaller pavilions: pretty Italian, Dutch and Swiss Cottages, Blank’s Hermitage and the old Orangery, where the State Ceramics Museum is located now, an extensive and absorbing collection of porcelain from the 18th century to the present day. On the other side of the lake is a large wood popular with local cyclists and joggers.

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  1. Mountain Lion Range

    mountain lion travel range

  2. A Trail Runner’s Guide to Mountain-Lion Encounters

    mountain lion travel range

  3. Mountain Lion Range

    mountain lion travel range

  4. Mountain Lion

    mountain lion travel range

  5. Mountain Lion Facts

    mountain lion travel range

  6. 10 Interesting Facts About Mountain Lions

    mountain lion travel range

VIDEO

  1. Mountain lion swims across river in Colorado

COMMENTS

  1. FAQ's

    Travel many miles at 10 mph; Reach speeds of 50 mph in a sprint; Identifying Mountain Lion Tracks. ... Though sizes vary greatly throughout the cat's geographic range, a typical adult male will weigh 110 to 180 pounds and the female 80 to 130 pounds. ... Mountain lions used to occupy the entire United States coast-to-coast, but today they are ...

  2. Mountain Lion

    Mountain lions can breed year-round. Female mountain lions usually give birth every two years. Litters can range in size from one to six cubs. The young may stay with their mother for as long as 26 months, but usually separate after about 15 months. In the wild, a mountain lion can live up to 10 years. In captivity, they can live up to 21 years.

  3. About Mountain Lions

    The mountain lion navigates its home range in a zigzag course, skirting open areas and taking advantage of available cover. The cat's keen senses are focused to pick up the slightest movement, odor, or sound. ... He found that the lions in his California study area travel at a steady speed of approximately one-half mile per hour. Though there ...

  4. Welcome to The Mountain Lion Tracking Webmap

    Welcome to The Mountain Lion Tracking Webmap A Project of UC Davis Wildlife Health Center & The Nature Conservancy. Esri, HERE, Garmin, FAO, USGS, NGA, ... Date Range. 13 6. 13 6. 12/4/2006 12/8/2011-+ Date Range: 10/31/2001 - 12/11/2016 Set date range and update lions. Time Options. Intervals: 5 Min: 1 Hr: 2 Hr: 6 Hr: Sync sunlight: Yes: No ...

  5. Cougar

    The cougar (Puma concolor) (/ ˈ k uː ɡ ər /, KOO-gər), also known as the panther, mountain lion, catamount and puma, is a large cat native to the Americas. It inhabits North, Central and South America, making it the most widely distributed wild, terrestrial mammal in the Western Hemisphere, and one of the most widespread in the world.Its range spans the Canadian Territory of Yukon ...

  6. Mountain Lion

    The Mountain Lion will return to feed on the prey for up to 10 days. The range of a Mountain Lion may cover 25 to 785 square miles. Here in southern Utah, lions have been known to occupy home ranges as large as 513 square miles. The size of a lion's territory depends on the availability of food and habitat quality.

  7. Mountain Lion

    A mountain lion walking away from an elk carcass in the woods. The mountain lion, also known as cougar, panther, puma, or catamount, is the one of the largest cats in North America and a top predator native to Valles Caldera. As part of predator-removal campaigns in the early 1900s, mountain lions, coyotes, and wolves were killed throughout the ...

  8. Mountain Lion

    Identification. They are the largest completely carnivorous animals in the park. At the shoulders, an adult mountain lion stands about 2-3 feet tall and 6 -7.5 feet long from the tip of the nose to the end of the tail. Mountain Lions have a round head and erect ears. They have powerful forequarters and necks, and long hind legs for leaping and ...

  9. Cougar Facts

    Cougars, also known as mountain lions or pumas, are native to Washington state. About 2,500 live in this state, double the populations of the early 1980s, and their numbers are growing. ... An adult cougar can range in length from 42 to 54 inches, with a 3-foot-long tail. Adult males weigh up to 200 pounds, and adult females up to 120 pounds. ...

  10. A Tale of Two Big Cats: What We Learned About the Mountain Lions in

    Mountain lions travel great distances and breed throughout their range, so gene frequencies throughout a particular state are likely similar. For now, it seems that this mountain lion's origin story will remain a mystery. Unlike the first mountain lion, the origin of the second mountain lion that moved through Illinois last fall is known.

  11. Mountain Lion

    By the late 1960s, mountain lions were given game-mammal status in many states, leading to a broader range of population management and programs. After this was enacted, their populations began steadily increasing in numbers. ... Due to the mountain lion's ability to travel long distances, the cats occasionally wander into areas that may seem ...

  12. 10 Interesting Facts About Mountain Lions

    Mountain lions are often considered umbrella species because of the large amounts of open habitat and wilderness they require to thrive. ... males have a home range covering from 50-150 square miles. Females have slightly smaller ranges of about 50 square miles. ... where young males may travel very far to establish their own area.

  13. Mountain Lion Population in Each State (Current Estimates)

    Mountain lions are found regularly throughout Idaho, and have been observed in nearly every county. Estimates place the mountain lion population in Idaho to be around 2,ooo . It is legal to hunt mountain lion in Idaho. In fact, in a controversial move, the state removed hunting quotas for mountain lions state wide.

  14. Cougar

    Cougar (Puma concolor) Cougar. (Puma concolor) Sleek and graceful, cougars ( Puma concolor ) are solitary and secretive animals rarely seen in the wild. Also known as mountain lions or pumas, cougars are known for their strength, agility, and awesome ability to jump. WDFW estimates there are approximately 3,600 cougars in Washington state.

  15. Mountain Lions in California

    Mountain lions are the second largest felid in North America with the largest geographic range of carnivores in the western hemisphere that stretches from southern Chile to the Canadian Yukon. The mountain lion is known by over 40 different common names including but not limited to puma, cougar, panther, red tiger, catamount, and screamer.

  16. Cougars in Minnesota

    The cougar - sometimes referred to as a mountain lion or puma - was found throughout most of Minnesota prior to European settlement, though never in large numbers. ... U.S. Cougar Range. Click to enlarge. Several hundred miles separate Minnesota from the nearest known self-sustaining breeding population of cougars, estimated to number ...

  17. Mountain Lion Research in Grand Canyon

    The mountain lion research conducted by Eric York prior to his tragic death in November 2007 has significantly added to our knowledge of this keystone predator in Grand Canyon National Park. ... A total of 127 traps were set along game trails, ridge lines, canyon walls and other common lion travel routes. At any one time, up to 25 traps were ...

  18. How California is helping its mountain lion population

    The 101 wildlife crossing project started because a lone mountain lion, known as P-22, would often roam Griffith Park, but was unable to cross the 101 to find a mate, CNN reported back in 2022. ...

  19. TPWD warn of mountain lion sighting in Trans-Pecos region

    The Texas Parks and Wildlife Department is warning residents in the Trans-Pecos region of reports of a mountain lion within city limits. ... 13 travel gadgets under $25 that will save your summer ...

  20. Mountain lion seen wandering West Texas town causes panic

    The Presidio Police Department said it first received a call about a puma wandering around town on April 9. Two responding police officers and animal control found the puma hiding in a stretch ...

  21. 107 Animals That Start With M

    Mountain Lion. This cat lives in mountain ranges across North America and is a relative of both cougars and bobcats. Mountain lions have been known to kill humans, though they typically avoid people and are more likely to try to avoid them than attack them. ... They tend to travel by running rather than walking to move quickly through their ...

  22. Moscow, Idaho

    First United Methodist Church (1904), S. Adams at E. 3rd St. Moscow (/ ˈ m ɒ s k oʊ / MOS-koh) is a city and the county seat of Latah County, Idaho.Located in the North Central region of the state along the border with Washington, it had a population of 25,435 at the 2020 census. Moscow is the home of the University of Idaho, the state's land-grant institution and primary research university.

  23. Mountain Lions

    Mountain Lions are the largest predators currently in Rocky Mountain National Park. They are also known as pumas, cougars and panthers. They vary in size and weight, with males reaching up to 200 pounds and eight feet in length (one-third of their length is the tail). Females are typically smaller. Mountain lion fur is unspotted and tan-brown ...

  24. Moscow parks

    Losiny Ostrov. Losiny Ostrov (Elk Island Park) Losiny Ostrov (Elk Island Park) is located at the north of Moscow. It covers 22 km from the west to the east and 10 km from the north to the south and it's one of the most beautiful national parks in Moscow. Two rivers, Yausa and Pechorka begin here.

  25. 21 Things to Know Before You Go to Moscow

    1: Off-kilter genius at Delicatessen: Brain pâté with kefir butter and young radishes served mezze-style, and the caviar and tartare pizza. Head for Food City. You might think that calling Food City (Фуд Сити), an agriculture depot on the outskirts of Moscow, a "city" would be some kind of hyperbole. It is not.